Acervus Kanouse
Pap. Mich. Acad. Sci., 23: 149 (1938)
Type species: Acervus aurantiacus Kanouse 1938
Type designated by Kanouse (1938)
Diagnosis: Apotheciis caespitosis, sessilibus, vel substipitatis in sclerotiformibus radicibus, coriaceis, fixis centralibus vel subcentricis, coloratis; hypotheciis parenchymaticis; excipulis prosenchymaticis; sporis ellipsoideis, non-septatis, hyalinis; paraphysibus crassis, irregulariter auctis, coloratis. Species typica Acervus aurantiacus.
Family: Otideaceae
Acetabula Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 330 (1870)
Type species: Acetabula vulgaris Fuckel 1870
Type designated by Clements & Shear (1931)
Diagnosis: Cupulae sparsae, magnae, in stipitem crassum, sulcatum retrorsum attenuatae, carnosae, campanulatae, extus venosae laevesve. Asci longissimi, cylindracei, 8spori. Sporidia in asci superiori parte oblique monosticha, ovata vel oblongo-ovata, hyalina, continua, enucleata uninucleatave. Paraphyses filiformes, clavatae. Terrestres, plerumque vernales.
Synonym of Helvella
Adelphella Pfister, Matočec & I. Kušan
Mycol. Montenegr., 11: 14 (2009)
Type species: Peziza babingtonii Berk. & Broome 1851
Type designated by Pfister et al. (2009)
Diagnosis: Pachyella similaris sed gelatinosa copiosa interna, apothecia mina et pulvinata. Sporae cum duabus guttulis prominentibus guttulisque multis minutis. Apothecia carens stratum exterius pilorum valli.
Family: Pezizaceae
Ahmadea Aman, Khalid & Moncalvo
Stud. Fung., 5 (1): 454 (2020)
Type species: Ahmadea dalanensis Aman & Khalid 2020
Type designated by Aman, Khalid & Moncalvo (2020)
Diagnosis: Hypogeous ascomata, irregular in shape, thin excipulum, solid gleba lacking paraphyses, asci with eight spores (sometimes less), ascospores globose, ornamented with sharp spines with a broad base and blunt ends.
Family: Pezizaceae
Aleuria Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 325 (1870)
Type species: Aleuria aurantia (Pers.) Fuckel 1870
Type designated by Clements & Shear (1931)
Diagnosis: Cupulae gregariae, magnae, ampliato-campanulatae, extus farinosae, stipitatae subsessilesve, carnosae, disco concavo, marginato, colorato. Asci longissimi, cylindracei, 8spori. Sporidia oblique monosticha, oblongo-ovata, 1—2 guttulata, continua, hyalina, episporio granuloso. Paraphyses filiformes, coloratae. Autumnales. Terricolae.
Family: Pyronemataceae
Aleurina Massee
Bull. Misc. Inf. Kew, 1898: 131 (1898)
Type species: Aleurina tasmanica Massee 1898
Type designated by Masse (1898)
Family: Pyronemataceae
Current definition: Apothecium solitary to gregarious, turbinate to deep cupulate when young, discoid to flat at maturity, cupulate to deep cupulate when dry; sessile to subsessile; medium sized. Hymenium olivaceous, olivaceous brown, brown, or purplish brown when fresh; olivaceous brown, dark brown or dark purplish brown when dry. Receptacle brown to reddish brown when fresh, dark brown to dark purplish brown when dry, usually with pustules at the flanks near the margin, and with subhyaline, flexuous hyphae at the base, basal hyphae often enmenshing soil particles. Ectal excipulum of textura angularis, cells light brown, isodiametric to elongated polygonal, axes of cells usually perpendicular to the outer surface of apothecium, walls somewhat rigid, cells of the pustules brown, subglobose, thick-walled; medullary excipulum of textura intricata, hyphae subhyaline to pale brown, running more or less parallel to the outer surface; subhymenium well-developed or indistinguishable. Asci cylindrical, 4- or 8-spored, J- in Melzer''s Reagent, usually with crozier at base. Ascospores ellipsoid, hyaline to pale yellow, with rounded warts, conical warts, or spines on surface, mostly biguttulate, uniseriate, fairly thick-walled when young, walls becoming thinner at maturity. Paraphyses simple, subcylindrical, subclavate to subcapitate at apex, with brownish pigment, septate in lower portion, in some species with a strongly cyanophilic paraphysis cap which soon loosens and either remains inverted cupulate or becomes reflexed and discoid, remaining attached to the paraphysis apex only at its center. On sandy or mossy soil, leaves, wood, or duff.
Amylascus Trappe
Trans. Br. mycol. Soc., 57 (1): 89 (1971)
Type species: Amylascus herbertianus (J.W. Cribb) Trappe 1971
Type designated by Trappe (1971)
Diagnosis: Ascocarpi subglobosi, plicati. Excipulm externum verrucosum, cellulis inflatissimis. Gleba venis coloratis, cellulosis. Asci sporis 5–8, parietibus amyloideis, incrassatis. Sporae globosae, ornatae.
Family: Pezizaceae
Current definition: Ascocarp subglobose, plicate. Peridium red to blackish brown, verrucose, tomentose, with an outer layer of greatly inflated (up to 100 µm) cells. Gleba trame not differentiated into fertile and non-fertile areas but deeply penetrated by meandering, red, peridium-line veins. Asci 5- to 8-spored, ellipsoid, sessile or with a short, simple stem, at maturity the walls more or less 2 µm thick and faintly blue in Melzer''s reagent. Spores globose, echinulate, hyaline.
Anthracobia Boud.
Bull. Soc. mycol. Fr., 1: 106 (1885)
Type species: Peziza melaloma Alb. & Schwein. 1805
Type designated by Boudier (1885)
Diagnosis: Poils courts et obtus ; spores avec ou sans sporidioles. Espèces à spores lisses ; carbonicoles
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, first cupuliform then discoid or flattened, hymenium yellow-orange to dull orangish, with an external surface covered by superficial light umber brown to brown pseudo-hairs, i.e. brown clavate or obtuse elongated cells. Margin hairy. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura subglobulosa/angularis. Asci operculate, arising from croziers, inamyloid, 8-spored. Paraphyses slender, enlarged to capitate at the top, hyaline. Ascospores uniseriate, ellipsoid, hyaline, biguttulate, thick-walled, smooth. Species saprobic on burnt soil.
Asexual morph of Scytalidium type.
Antrelloides P.S. Catches. & D.E.A. Catches.
Swainsona, 31 (6): 82 (2018)
Type species: Antrelloides atroceracea P.S. Catches. & D.E.A. Catches. 2018
Type designated by Catcheside & Catcheside (2018)
Family: Pezizaceae
Current definition: Apothecial ascomata up to 40 mm diam. Hymenium convex; firm, waxy; black, violaceous-black. Basal stipe substantial; obconic; a chambered base of knobby columnar ridges radiating from base; fragile, waxy. Asci amyloid; cylindrical; operculate; 8-spored. Ascospores ellipsoid; smooth under light microscope. Paraphyses longer than asci; septate; tips swollen and tending to aggregate, encrusted with brown amorphous matter; branching occasionally near tips. Medullary excipulum of several layers of interwoven hyphae, grading into ectal excipulum. Ectal excipulum of large globose to subglobose cells with interwoven hyphae. Stipe tissue of textura globulosa and chains of subglobose cells.
Aparaphysaria Speg.
Anal. Soc. cient. argent., 94 (1-2): 81 (1922)
Type species: Aparaphysaria doelloi Speg. 1922
Type designated by Spegazzini (1922)
Diagnosis: Char. Pezizea, subceraceo-carnosa, glaberrima, hyalospora; ascomata mediocra subturbinato-cupulata, sessilia; asci cylindracei densissime constipati, octospori, operculatim dehiscentes, paraphysibus omnino nullis; sporae ellipticae, continuae, hyalinae, laeves.
Est Peziza paraphysibus plane destituta.
Family: Pyronemataceae
Number of species: 2 fide DoF
Aquapeziza D.M. Hu, L. Cai & K.D. Hyde
Mycologia, 104 (2): 541 (2012)
Type species: Aquapeziza globispora D.M. Hu, L. Cai & K.D. Hyde 2012
Type designated by Hu et al. (2012)
Family: Pezizaceae
Number of species: 1
Current definition: Apothecia scattered, epigeous, sessile, turbinate, white; excipulum composed of an ectal exipulum of textura globulosa to textura angularis and medullary excipulum of textura intricata. Paraphyses cylindrical, septate, hyaline. Asci unitunicate, not protruding, operculum not distinct, at first broad cylindrical becoming ovoid at maturity, with a rounded apex, hyaline, amyloid, eight-ascospored. Ascospores globose, one-celled, guttulate, hyaline, smooth.
Arpinia Berthet
Bull. mens. Soc. linn. Lyon, 43, num. spéc.: 36 (1974)
Type species: Arpinia inops Berthet 1974
Type designated by Berthet (1974)
Diagnosis: Apotheciae terrestres, pedunculatae, cupulares, extrinsecus subtiliter furfuraceae, omnino pallidae, pigmentis carotenoidibus indigentes. Asci octospori, non iodo caerulescentes. Sporae uni – aut biguttulatae, ovatae, laeves, uninucleatae. Paraphyses clavatae, apice rectae. Ascogenae hyphae fibulatae.
Family: Otideaceae
Ascobolus Pers.
Syst. Nat., Edn 13, 2 (2): 1461 (1792)
Type species: Ascobolus pezizoides Pers. 1797 (= Peziza stercoraria Bull.)
Type designated by Persoon (1792)
Diagnosis: Fungus hemisphaericus: Vesicula oblonga diseo sparsim subimmersa elastice ejiciens femina.
Family: Ascobolaceae
Current definition: Apothecia superficial or immersed, sometimes with a short stalk; mostly small, 0.3-10 mm, in one species up to 30 mm diameter. Receptacle subglobular, pyriform, obconical, cup- or saucer-shaped, rarely lenticular; surface smooth, furfuraceous villose or downy. Asci saccate-clavate or cylindric-clavate; at maturity the tips strongly protruding above the hymenium; with a rounded, dome-shaped or slightly truncate apex. Ascospores biseriate or irregularly disposed; free, neither clustered nor arranged according to a regular pattern (as illustrated in fig. 00) during any phase of the maturation. Paraphyses slender, cylindrical; often embedded in colourless, yellowish or yellowish-green mucus.
Predominantly fimicolous, but also foliicolous, lignicolous, terrestrial and pyrophilous.
Asexual morph of Rhizostilbella type.
Ascocalathium Eidam ex J. Schröt.
in Cohn, Krypt.-Fl. Schlesien, 3.2 (1-2): 32 (1893)
Type species: Ascocalathium stipitatum Eidam ex J. Schröt. 1893
Type designated by Schröter (in Cohn, 1893)
Diagnosis: Fruchtkörper frei aufsitzend, sehr klein, scheibenförmig aus kleinen fleischigen Grundlagen un den frei auf ihnen aufsitzenden Schläuchen mit dazwischen stehenden Paraphysen gebildet. Schläuche cylinderisch, 8sporig. Sporen ellipsoidisch; Membran farblos. Paraphysen haarförmig, die Schlaüche überragend.
Family: Pyronemataceae
Number of species: 1 fide DoF
Ascodesmis Tiegh.
Bull. Soc. bot. Fr., 23: 275 (1876)
Type species: Ascodesmis nigricans Tiegh. 1876
Type designated by Seaver (1928)
Diagnosis: Comme l''exprime le nom d''Ascodesmis, que je propose de leur donner, le fruit mûre ces plantes se compose simplement d''un bouquet ou d''une rosette d''asques divergents, entremêlés de paraphyses, le tout inséré à la surface supérieure d''une petite masse cellulaire arrondie, qui à son tour s''attache au milieu de sa face inférieure et par un court rameau à un filament mycélien. Complétement dépourvus d''enveloppe ou de périasque, ces fruits sont très petits, les plus gros atteignant à peine 2 à 3 dixièmes de millimètre; aussi n''apparaissent-ils à l''oeil nu, sur la trame blanche formée par le mycélium aérien de la plante, que comme autant de petils points d''un brun-chocolat dans l''Ascodesmis nigricans, d''un jaune d''or dans l''Ascodesmis aurea. […]
Family: Ascodesmidaceae
Ascophanus Boud.
Ann. Sci. Nat., Bot., sér. 5, 10: 241 (1869)
Type species: Ascobolus granuliformis P. Crouan & H. Crouan 1858
Diagnosis: Ascobolus et Peziza auctorum.
Receptaculum carnosum, glabrum, aut pruinosum, aut etiam pilosum, rario marginatum, semper sessile. Hymenium planum aut convexum, gelatina raro conspicua, thecis parum prominentibus, semper crystallino-papillatum. Paraphyses thecarum longitudine, simplices aut ramosae, septatae, graciles aut ad apicem incrassatae, hyalinae aut coloratae, intus saepius granulosae. Thecae crassae clavatae, raro oblongo-ovatae, hyalinae aut vix tinctae, ad basin attenuatae, sporas 8 includentes (unica specie thecas 16 sporas offert), operculo conspicuo, rotundato, saepius recurvato, dehiscentes. Sporae ovatae aut oblongo-ovatae, semper hyalinae aut vix tinctae, nudae sed juniores gelatina singulatim circumdatae, episporio hyalino, levi, raro minutissime punctato, nucleo conspicuo ; maturae ad extremitatem thecae sic aggregatae : sex in duabus seriebus juxtapositis et una utraque extremitate.
Synonym of Iodophanus ?
Ascorhizoctonia Chin S. Yang & Korf
Mycotaxon, 23: 468 (1985)
Type species: Ascorhizoctonia praecox Chin S. Yang & Korf 1985
Type designated by Yang & Korf (1985)
Diagnosis: Coloniae celeriter crescentes, hyalinae vel brunnae, patellas petrianas 90 mm latas agarum + aquam, agarum + Zeae farinam, agarum + hordei tosti extractum, agarumve + Solani tuberosi dextrosum continentes 5–7 diebus temperatura ca 21 C velantes. Mycelium adpressum, immersum, superficiareve, annulis concentricis praesentibus absentibusve, cellularum monilioidearum caespites sporodochiis similes super agaro vel intra agarum saepe producentibus. Hyphae cylindrales, septatae, 2.0–13.0 mm latae, hyalinae vel brunneae, laeves vel sparsim verrucosae, aut parallele jacentes aut paene ad angulum 90° ramificantes, cellulis globulis oleosis repletis. Circuli et ponticuli hyphales praesentes vel absentes. Nec sclerotia vera nec bulbili procreati.
Family: Pyronemataceae
Current definition: Apothecia sunken in the substratum up to halfway up the apothecial flanks; ascospores ellipsoid with conspicuously tapered ends, walls thick (0.7–1 μm), perispore cyanophilic, not loosening in CB, smooth or finely verrucose, RBI 1–2, containing only BSG; asci arising from compact croziers; hairs distributed only on marginal area, hair base narrow; colonies concentrically banded on MEA, of moderate growth (reaching 5–6 cm diam. on MEA at fourth day); ascorhizoctonia-type anamorph normally formed on MEA; mycelia forming loops on water agar and CMA; saprotrophic; so far including species with yellowish to ochre hymenia in fresh apothecia, occurring only on burnt substrates.
Ascosparassis Kobayasi
Bull. nat. Sci. Mus., Tokyo, n.s., 5: 45 (1960)
Type species: Ascosparassis shimizuensis Kobayasi 1960
Type designated by Kobayasi (1960)
Diagnosis: Fructificationes e sclerotio subterraneo carnoso oriundae, stipitatae coriaceae flabellatim dilatatae, margine lobatae undulatae integrae; hymenio unilaterali laevi; excipulo externo verrucoso parenchymatico, hypothecio prosenchymatico. Paraphyses filiformes simplices, apice hamatae; ascis cylindricis octosporis inoperculatis J-; ascosporis uniseriatis ellipsoideis continuis laevibus hyalinis.
Family: Otideaceae
Current definition: Fructifications growing from underground sclerotioid carnose mycelial mass, stipitate coriaceous, flabellately dilated, margin lobed, undulate, entire; hymenium unilateral smooth; exciple (ectal excipulum) verrucose, parenchymatic; inner tissue (medullary excipulum) prosenchymatic. Paraphyses simple, not forming epithecium, apex peg-shaped; asci cylindric 8-spored inoperculate, apex opening by pore, not bluing with iodine. Ascospores uniseriate, ellipsoid smooth one-celled, hyaline.
Aurantiolachnea Van Vooren
Ascomycete.org, 13 (1): 9 (2021)
Type species: Lachnea solsequia Quél. 1892
Type designated by Van Vooren et al. (2021)
Diagnosis: Differs from Humaria by the orange-coloured hymenium, smooth or finely dotted ascospores, its saprobic trophic status and genetic profile.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, orange-coloured, with an external surface and margin covered by brown hairs. Excipulum two-layered: medullary layer of textura intricata, ectal layer of textura subglobulosa/angularis. Hairs simple, superficial, septate, with a simple base. Asci operculate, narrowing towards the base, arising from croziers, inamyloid, 8-spored. Paraphyses slender, not enlarged at the top, containing carotenoid pigments. Ascospores uniseriate, ellipsoid, hyaline, biguttulate, smooth or very finely dotted. Trophic status probably saprobic. Asexual morph unknown.
Aurophora Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 52 (1968)
Type species: Peziza dochmia Berk. & M.A. Curtis 1868
Type designated by Rifai (1968)
Diagnosis: Apothecia gregaria, magna, stipitata vel subsessilia, inaequalia, flabelliformis, auriformis vel rara discoidea. Caro (excipulum medullare) e textura intricata, hyphis septatis, gracilibus, gelatinosis composita. Asci subcylindracei, inferne longe angustati, octospori. Ascospori uniseriati, hyalini, guttulati, leniter curvati, striati. Paraphyses lineares, septatae, ramosae. Hab. ad lignum putridum.
Family: Sarcoscyphaceae
Number of species: 1 fide DoF
Babosia D.G. Knapp, Zagyva, Trappe & Kovács
Mycologia, 112 (4): 811 (2020)
Type species: Babosia variospora D.G. Knapp, Zagyva, Trappe & Kovács 2020
Type designated by Knapp et al. (2020)
Diagnosis: Ascomata gasteroid, hypogeous. Differs from the phylogenetically most closely related Temperantia, Hydnobolites, and Stouffera by the dark sporogenous zone and dark brown spore color at maturity, the clear peridermal layer at maturity, and the variability of the ornamentation of the ascospores.
Family: Pezizaceae
Number of species: 1
Balsamia Vittad.
Monogr. Tuberac.: 30 (1831)
Type species: Balsamia vulgaris Vittad. 1831
Type designated by Clements & Shear (1931)
Diagnosis: Char. Uterus mollis, sessilis, arrhizus, semper clausus, celluloso-carnosus. Sporangia oblonga, membranacea, pedicellata, octospora, cellularum parietibus immersa ac seriatim disposita. Sporidia cylindracea, laevia, pellucida.
Family: Helvellaceae
Barssia Gilkey
Mycologia, 17 (6): 253 (1925)
Type species: Barssia oregonensis Gilkey 1925
Type designated by Gilkey (1925)
Diagnosis: Ascocarp scabrous to verrucose, reddish-yellow, nearly even to lobed, somewhat flattened with an irregular opening at the apex into a central depression; cortical structure of the surface of the ascocarp carried into the depression except where the hymenium projects inot it; inner tissue of the ascocarp thrown up in more or less connected folds, forming canals and chambers lined with hymenium, these canals opening into the cavity of the ascocarp; hymenium composed of regularly arranged asci and paraphyses; paraphyses very slender and much longer than the asci; asci cylindrical, somewhat club-shaped, 8-spored; spores ellipsoid, smooth, 1- or incompletely 2-seriate, colorless.
Family: Helvellaceae
Berggrenia Cooke
Grevillea, 8 (45): 64 (1879)
Type species: Berggrenia aurantiaca Cooke 1879
Type designated by Cooke (1879)
Diagnosis: Carnosa, inflata, subsessilis, intus hymenio effuso obducta. Sporidiis amplis, ascigeris. Gard. Chron., Oct., 1879
Family: Incertae sedis
Boubovia Svrček
Česká Mykol., 31 (2): 71 (1977)
Type species: Humaria luteola Velen. 1934
Type designated by Svrček (1977)
Diagnosis: Apothecia minuta vel mediocria, sessilia, concava, discina, crasse carnosa, margine integro, nuda, luteo-colorata. Excipulum ectale medullareque textura globulosa vel subglobulosa, cellulis ecoloratis, parietibus haud incrassatis. Asci cylindracei, non amyloidei, octospori. Paraphyses tenuiter filiformes, apice non dilatatae, conspecte curvatae et spiraliter contortae. Ascosporae ellipsoideae, verrucosae, hyalinae. Hab. ad terram.
Family: Coprotaceae
Boudiera Cooke
Grevillea, 6 (38): 76 (1877)
Type species: Boudiera areolata Cooke & W. Phillips 1877
Type designated by Boudier (1885)
Diagnosis: Immarginata, orbiculare, discoidea, plana, vel convexa, carnosa; hymenio papillato; ascis clavatis vel clavato-cylindraceis, exclusis; sporidiis globosis; paraphysibus distinctis.
Family: Pezizaceae
Boudierella Sacc. ex Marchal
Bull. Soc. Roy. Bot. Belg., 34: 130 (1895)
Type species: Boudierella cana Marchal
Type designated by Marchal (1895)
Diagnosis: Ascomata carnosula, sessilia, hemisphaerica. Discus planus, margine ciliatus, ascis papillatus. Asci oblongi, 4-spori, fissura longitrorsus dehiscentes, paraphysati. Sporidia globosa hyalina, laciniis numerosis, angustis, insigniter contecta.
A gen. affini Boudiera differt sporidiis non octonis nec coloratis ascisque longitrorsus dehiscentibus.
Family: Pyronemataceae
Number of species: 1 fide DoF
Byssonectria P. Karst.
Meddn Soc. Fauna Flora fenn., 6: 6 (1881)
Type species: Byssonectria obducens P. Karst. 1881
Type designated by Karsten (1881)
Diagnosis: Perithecia stromate byssino, effuso, connexa, ceteroqvin, sicut etiam asci et sporae, Nectriellae; paraphyses numerosae.
Family: Pyronemataceae
Caccobius Kimbr.
Am. J. Bot., 54 (1): 22 (1967)
Type species: Caccobius minusculus Kimbr. 1967
Type designated by Kimbrough & Korf (1967)
Diagnosis: Apothecia minuta, 150-175 µ lata, pallida vel alba, solitaria vel se congregantia, sessilia, discoidea; excipulum basale, margine setis septatis 24-40 µ longis, 2.5-3.0 µ latis instructa; asci 15-25, cylindrici, 125-135 × 50-60 µ, in iodo non caerulescentes, in "congo red" aequabiliter tincti epistomio apicali hyalino 15-20 µ diam. excepto quod atramento "Waterman''s blue-black" tingi potest; ascospori elliptici, 4.9-5.2 × 2.1-2.3 µ, muris usque ad 0.5 µ crassis, sine vacuolis evidentibus; paraphyses multae, filiformes, septatae, in tunica mucilaginea ascum quemque involvente mersae, pseudo-excipulum hypharum parallelarum rectarum supra excipulum verum formantes, epithecium quoque supra ascum quemque ramis efficientes.
Family: incertae sedis
Number of species: 1 fide DoF
Caloscypha Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Peziza fulgens Pers. 1822
Type designated by Boudier (1885)
Diagnosis: Thèques de grandeurs ordinaires ou petites (brévithèques). Paraphyses rameuses, souvent atténuées aux extrémités ; thèques petites ; spores rondes.
Family: Caloscyphaceae
Asexual morph of Geniculodendron type.
Carbomyces Gilkey
N. Amer. Fl. Ser., 2, 1: 27 (1954)
Type species: Carbomyces emergens Gilkey 1954
Type designated by Gilkey (1954)
Diagnosis: Ascocarp compact, hypogaeous or epigaeous, more or less globose but with a suggestion of a slightly narrowed, darker base; peridium firm when dry, its texture somewhat that of carbonized wood; outer layer coarsely pseudoparenchymatous; innermost layer of coarse, free or more or less fused, branching hyphae, this layer continuing into the gleba as a network of veins surrounding nests of closely-packed minute asci; mature asci brownish, broadly obovate to globose, thick-walled, often eventually released from the hyphae, and tardily and irregularly dehiscent to allow gradual escape of the spores; spores 8, globose or broadly ellipsoid, smooth or with minute scattered papillae, brownish.
Family: Pezizaceae
Cazia Trappe
Mem. N. Y. bot. Gdn, 49: 336 (1989)
Type species: Cazia flexiascus Trappe 1989
Type designated by Trappe (1989)
Family: Pezizaceae
Cephaliophora Thaxt.
Bot. Gaz., 35: 157 (1903)
Type species: Cephaliophora tropica Thaxt. 1903
Diagnosis: Vegetative hyphae copious, branching, septate, colorless. Sporophores arising as short branches from the hyphae, which become more or less abruptly enlarged distally to form a variably differentiated head, from the surface of which the spores are produced. Spores once to several times transversely septate, becoming brownish, the sterile basal segment narrowed to form a more or less distinct pedicellate attachment.
Family: Ascodesmidaceae
Number of species: 6 fide DoF
Chaetothiersia B.A. Perry & Pfister
Fungal Divers., 28: 69 (2008)
Type species: Chaetothiersia vernalis B.A. Perry & Pfister 2008
Type designated by Perry & Pfister (2008)
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, cupuliform, pale grey to whitish-coloured or yellow to orangish yellow, with an external surface covered by brown hairs. Margin hairy. Excipulum two-layered: medullary layer thick, of textura intricata, and ectal layer of textura globulosa/angularis. Hairs superficial, simple, septate, with a simple ± swollen base. Asci operculate, narrowing toward base, arising from free croziers, inamyloid, 8-spored. Paraphyses slender, equal to enlarged (sometimes subcapitate) at the top, hyaline or containing a sparse granular yellow pigment. Ascospores uniseriate, ellipsoid, hyaline, guttulate or not, smooth. Species saprobic on the litter and woody debris of conifers. Asexual morph unknown.
Chalazion Dissing & Sivertsen
Norw. J. Bot., 22 (1): 1 (1975)
Type species: Chalazion sociabile Dissing & Sivertsen 1975
Type designated by Dissing & Sivertsen (1975)
Diagnosis: Carposoma parvum, superficiale, discoides vel pulvinatum, albidum vel colore perpallido, grandinem parvam revocans, unde nomen. Excipulum exterius simplex, e cellulis cyanophilis membranis tenuibus compositum, nullis pilis vestitum. Asci operculati, magni, prominentes, non amyloides, clavati, membranis crassis. Paraphyses hyalinae vel colore perpallido, septatae, sursum incrassatae. Ascosporae biseriatae, membranis crassis, colore nullo, maturae ornamentis cyanophilis notatae, guttulas continentes.
Family: Ascodesmidaceae
Current definition: Fruit bodies small, superficial, discoid to pulvinate, whitish or very lightly pigmented. Outer excipullum simple, of thin walled, cyanophilic cells, without hairs. Asci operculate, large, protruding, non-amyloid, clavate, thick walled. Paraphyses hyaline or very lightly coloured, enlarged above, septate. Ascospores thick walled, biseriate, colourless, with guttules, at maturity with a cyanophilic ornamentation.
Cheilymenia Boud.
Bull. Soc. mycol. Fr., 1: 105 (1885)
Type species: Peziza stercorea Pers. 1800
Type designated by Clements & Shear (1931)
Diagnosis: Poils du réceptacle longs et aigus. Hyménium rouge ou jaune, rarement de couleur fauve ou pâle. Spores n''ayant jamais intérieurement de gouttelettes. Hyménium jaune ou orangé. Espèces fimicoles.
Family: Pyronemataceae
Choiromyces Vittad.
Monogr. Tuberac.: 50 (1831)
Type species: Choiromyces meandriformis Vittad. 1831
Type designated by Clements & Shear (1931)
Diagnosis: Char. Uterus polymorphus, sessilis, arrhizus, extus laevis, interne carnosus, solidus, caro albida, venis proprie seminiferis, decolorantibus variegata. Sporangia longissime pedicellata, lagenaeformia, octospora, simplici serie ad latera venarum distributa. Sporidia sphaerica.
Family: Tuberaceae
Chorioactis Kupfer ex Eckblad
Nytt Mag. Bot., 15 (1-2): 127 (1968)
Type species: Urnula geaster Peck 1894
Type designated by Eckblad (1968)
Diagnosis: Apothecia juvenilia clausa, matura ex apice basin versus stellatim dehiscentia, excipulo toto textura intricata constructo, extus tomentosa, tomento e stratis cellularibus externis formato; pili tomenti simplices, septati, recti, pariete duplici fusco instructi. Asci cylindrici, non amyloidei, octospori. Sporae fusiformes, saepe asymmetricae, laeves, hyalinae. Cellulae paraphysium ante ejaculationem sporarum distincte turgescentes.
Family: Chorioactidaceae
Current definition: Apothecia closed when young, splitting stellately from the apex downwards when ripe. Externally tomentose. Excipulum of a textura intricata throughout. The outer cell layers give rise to the tomentum which consists of a single type of hair. Hairs unbranched, septate, straight, and with a brown, double wall. Asci cylindrical, eight-spored, non-amyloid. Spores fusiform, smooth, hyaline. Paraphyses with cells that swell considerably before spore discharge.
Chromelosporium Corda
in Sturm, Deutschl. Fl., 3 Abt., 3: 81 (1833)
Type species: Chromelosporium ochraceum Corda 1833
Type designated by Corda, in Sturm (1833)
Diagnosis: Char. gen. Sporae continuae coloratae, in gelatina nidulantes, floccis heterogeneis destructis ramosis, articulatis hyalinis inspersae. Acervuli effusi colorati.
Family: Pezizaceae
Number of species: 3 fide DoF, currently 5
Cidaris Fr.
Summa veg. Scand., sectio post.: 347 (1849)
Type species: Verpa caroliniana Schwein. 1832
Type designated by Cooke (1875)
Diagnosis: Verpa Schwein.! Discus pileatus, rugoso-tuberculosus, margine revoluto. Stipes rugoso-plicatus.
Family: Helvellaceae
Number of species: 1 fide DoF
Ciliaria Quél. ex Boud., nom. illeg.
Bull. Soc. mycol. Fr., 1: 105 (1880)
Type species: Peziza scutellata L. 1753
Type designated by Clements & Shear (1931)
Diagnosis: Poils du réceptacle longs et aigus. Spores lisses ou plus souvent verruqueuses, garnies intérieurement de gouttelettes nombreuses, très rarement presque nulles. Hyménium rouge, espèces terrestres ou humicoles.
Synonym of Scutellinia
Cleistoiodophanus J.L. Bezerra & Kimbr.
Am. J. Bot., 63: 838 (1976)
Type species: Cleistoiodophanus conglutinatus J.L. Bezerra & Kimbr. 1976
Type designated by Bezerra & Kimbrough (1976)
Diagnosis: Apothecia coprophila, gregaria vel cespitosa, sese cleistohimenialiter evolventia, occulsa, autem, manentia, albo colore praedita; excipula ectalis cyanophilica, cellulas rectangulares vel quadratas habens; excipula medularis cyanophilica, cum textura globulosa; asci generaliter octospore, diffuse amiloides, large clavati, cum parietibus crassis in juventute, pedicelati; ascospori uninucleati, hyalini, ellipsoides, laeves, perisporium mucilaginosum ostentantes; paraphyses ramosae, hyalinae, septatae, multinucleatae.
Momentum conidiale: Oedocephalum Preuss.
Family: Ascobolaceae
Current definition: Apothecia gregarious to cespitose, developing cleistohymenially, remaining closed, light colored; ectal excipulum of textura prismatica, cyanophilic, of rectangular or quadratic cells; medullary excipulum cyanophilic, of textura globulosa; asci usually 8-spored, diffusely amyloid, broadly clavate, thick-walled in youth, stalked; ascospores uninucleate, hyaline, ellipsoid, smooth, a mucilaginous perispore present; paraphyses branched, hyaline, septate, multinucleate; conidial stage: An Oedocephalum Preuss. Coprophilous.
Cleistothelebolus Malloch & Cain
Can. J. Bot., 49 (6): 851 (1971)
Type species: Cleistothelebolus nipigonensis Malloch & Cain 1971
Type designated by Malloch & Cain (1971)
Diagnosis: Ascogonia convoluta. Ascocarpae ab subglobosis irregulares, pallide brunneae, nonstromaticae. Asci irregulariter dispositi, ovoidei, evanescentes. Ascosporae unicellares, hyalinae, sine pora, sine "de Bary bubbles." Conidia modo blastosporae producta, hyalina, unicellaria, frequenter gemmifera.
Family: Pyronemataceae
Number of species: 1 fide DoF
Current definition: Ascocarp initials coiled. Ascocarps subglobose to irregular in shape, pale brown, nonostiolate, nonstromatic. Asci irregularly disposed, ovoid, evanescent. Ascospores one-celled, hyaline, without germ pores, without "de Bary bubbles." Conidia produced as blastospores, hyaline, onecelled, often budding.
Complexipes C. Walker
Mycotaxon, 10 (1): 99 (1979)
Type species: Complexipes moniliformis C. Walker 1979
Type designated by Walker (1979)
Diagnosis: Sporocarpia ignota. Chlamydosporae globosae, 55-110 µm in diam, ferruginae vel atrocinnamomeae, tunica stratis tribus: exteriore ± 1 µm, medio usque ad 6 µm, interiore 0.5-1 µm. Pagina sporae rugosa vel papillosa, altitudo ornamenti usque ad 4 µm. Hypha affixa papillosa vel interdum rugosa, bitunicata, septata cellusis usque ad novem, ad septa constricta, cellula ad basim sporae cupulata, usque ad 24 × 24 µm. Typus: Ames, Iowa, U.S.A., Walker #27 (OSC).
Family: Pyronemataceae
Number of species: 2 fide DoF
Conoplea Pers.
Tent. disp. meth. fung.: 55 (1797)
Type species: Dematium sphaericum Pers. 1795
Diagnosis: Fila persistentia in formam distinctam compacta, intus pulvere referta.
Family: Sarcosomataceae
Number of species: 8 fide DoF
Cookeina Kuntze
Revis. gen. pl., 2: 849 (1891)
Type species: Peziza tricholoma Mont. 1834
Type designated by Seaver (1927)
Diagnosis: Cookeina OK. = Trichoscypha Cooke 1879 non Hk.f.* in BHgp. 1862. Die altere Anacardiacee gilt, sodass die Discomycetee anders zu benennen ist; sie moge den berühmten Namen ihres Begrunders tragen. Die Arten sind nach Saccardo sylloge VIII 160/3 übertragen: Cookeina [...]
Family: Sarcoscyphaceae
Coprobia Boud.
Bull. Soc. mycol. Fr., 1: 107 (1885)
Type species: Peziza granulata Bull. 1790
Type designated by Boudier (1885)
Diagnosis: Réceptacles marginés, à marge dépassant ou ne d"passant pas l''hyménium, mais toujours l''entourant sur les côtés. Spores sans sporidioles, lisses ; espèces furfuracées, stercoricoles.
Synonym of Cheilymenia
Coprotus Korf & Kimbr.
Am. J. Bot., 54 (1): 21 (1967)
Type species: Ascobolus sexdecimsporus P. Crouan & H. Crouan 1858
Type designated by Kimbrough & Korf (1967)
Diagnosis: Apothecia sessilia, discoidea vel aliquantulum convexa, solitaria vel se congregantia, plerumque quam 1 mm diam, minora, ascis prominentibus asperata; asci operculati, 30 vel plures, 30-200 µ longi, 5-60 µ lati, in iodo non caerulescentes, in "congo red" aequabiliter tincti; ascospori in 1 vel 2 seriebus vel irregulariter dispositi, 8-256 in asco quoque, leves, elliptici, 6.0-16 × 3.5-10 µ, muris tenuibus, bullulam conspicuam continentes; paraphyses filiformes, septatae, uncinatae vel clavatae, simplices vel saepe ramosae.
Family: Coprotaceae
Cubonia Sacc.
Syll. fung., 8: 527 (1889)
Type species: Lasiobolus brachyascus Marchal 1885
Type designated by Seaver (1927)
Diagnosis: Lasioboli et Ascophani spec. Auct. — Ascomata sessilia hemisphaerico-obconica, glabra vel pilosa, disco plano, ascis papillato. Asci clavati vel subpiriformes, octospori, paraphysati. Sporidia perfecte sphaerica, asperula, hyalina.
Synonym of Ascobolus
Cupulina Dougoud, Van Vooren & M. Vega
Ascomycete.org, 7 (2): 39 (2015)
Type species: Cupulina montana Dougoud, Van Vooren & M. Vega 2015
Type designated by Dougoud et al. (2015)
Family: Pyronemataceae
Current definition: Apothecia gregarious, hemispherical or cupulate, sessile, partially embedded in the substrate or superficial, small, brownish ochraceous yellow, brownish orange or orange. Margin protruding, grossly and irregularly crenulate or fimbriate, whitish to pale yellow. Asci operculate, octospored, inamyloid. Ascospores fusoid or ellipsoid, hyaline. Paraphyses straight, enlarged at the top, with yellowish or hyaline content, without iodine reaction, septate, simple or furcate. Medullary excipulum of textura intricata. Ectal excipulum of textura globulosa to globulosa-angularis. Habitat: on soil, on naked or mossy ground.
Cyathipodia Boud.
Hist. Class. Discom. Eur.: 39 (1907)
Type species: Cyathipodia platypodia Boud. 1907
Type designated by Dissing (1966)
Diagnosis: Ce genre est bien voisin des Leptopodia dont il ne diffère que par les réceptacles qui ne sont jamais à bords infléchis mais toujours cupulaires. Le pédicule est long, grèle, ordinairement non sillonné ou à peine lacuneux, plus rarement sillonné. Les thèques, spores et paraphyses sont celles des Helvelles. Toutes les espèces sont terrestres.
Synonym of Helvella
Daleomyces Setch.
Mycologia, 16 (5): 240 (1924)
Type species: Daleomyces gardneri Setch. 1924
Type designated by Setchell (1924)
Diagnosis: Ascomata magna, inferne per stipitem et contextam mycelialem curtam robustamque, truncato obconicam, affixa, peridio maturitate proprio nullo vestita, interne labyrinthiformi-cavernosa; cameris numerosis cavis, clausis, forma magnitudineque irregularibus, earum parietibus hymenio vestitis, ascis elongatis, cylindraceis, octosporis, paraphysibus intermixtis; paraphysibus moderate robustis, ascos distincte superantibus, earum partibus exsertis flexis; sporis non septatis, oblongis, leviter ruguloso-reticulatis.
Family: Pezizaceae
Current definition: Ascomata epigeous, cupulate, discoid or sparassoid (cabbage-head), of various colours: whitish with pinkish tinges, becoming more or less brownish with age, purplish brown, olivaceous brown or distinctly violet. Flesh without latex. Asci operculate, 8-spored, with the top strongly bluing in an iodine solution (WTR type), with or without crozier. Paraphyses containing a vacuolar pigment, brown, purple brown or yellowish becoming brown with maturity. Ectal excipulum of textura globulosa, with large cells, sometimes partly gelatinised in the outer part (e.g. D. exogelatinosus). Ascospores biguttulate, ornamented with isolate warts, punctiform or irregular, often more dense at the poles. Species saprobic, preferentially growing on basic substrates (burned soil, sandy soil, soil with a high pH).
Delastria Tul. & C. Tul.
Ann. Sci. Nat. Bot., sér. 2, 19: 379 (1843)
Type species: Delastria rosea Tul. & C. Tul. 1843
Type designated by Tulasne & Tulasne (1843)
Diagnosis: Peridium tenuissimum byssaceum, candidum, arenae immistum; substantia carnosa, molliuscula, humida, in glebulis versiformibus septis mollibus albis immutabilibus (peridii filamentis stipatis intropulsis aequiparandis) dissecta ; sporangia ovato-oblonga, obtusa saepius incurvato-subreniformia, in glebulis nidulantia, sporas 3 (rarius 4) sphaericas reticulato-echinatas foventia ; guttula (?) in quaque spora subconcentrica. — Fungus basi absorbenti patentissima eradicata donatus.
Family: Pezizaceae
Dennisiopsis Subram. & Chandrash.
Kew Bull., 31 (3): 639 (1977)
Type species: Dennisiopsis octospora Subram. & Chandrash. 1977
Type designated by Subramanian & Chandrashekara (1977)
Diagnosis: Discomyces apothecia gerens. Apothecia superficialia, sparsa, sessilia, laete flava vel cremea, e fasciculo vel e fasciculis pluribus ascoram et paraphysium in medulla basili constata; medulla sine excipulo ectali. Asci operculati 8- vel multispori, non-amyloidei. Ascosporae hyalinae vel pallide flavae, unicellulares, tenuiter tunicatae, globosae vel ellipsoideae, laeves, in massam interdum agglutinantes, unaquaque ''bulla de Baryea'' prominenter ornata. Paraphyses filiformes, septatae, simplices vel ramosae.
Family: Incertae sedis
Number of species: 2 fide DoF
Densocarpa Gilkey
N. Amer. Fl., ser. 2, 1: 15 (1954)
Type species: Densocarpa shanorii Gilkey 1954
Type designated by Gilkey (1954)
Diagnosis: Ascomatibus hypogaeis aut epigaeis, compactis, puncto adjunctionis myceliali instructis, superficie glabris vel minute velutinosis, saepe tessellatis; texto corticis minute pseudoparenchymatico; gleba firmula lacunis multiformibus, saepe gyroso-labyrinthicis, floccis trajectis, hymenio ex ascis paraphysibusque in valium ordinatis vestitis; ascis cylindricis aut lente clavatis, octosporis; sporis globosis, levibus, saepe 1-4 tantum maturantibus, uniseriatis.
Family: Tarzettaceae
Current definition: Ascocarp hypogaeous or epigaeous, firm, with a broad mycelial area at the base, the upper surface often minutely downy and tending to crack irregularly, appearing tessellate at maturity; gleba penetrated by numerous chambers and winding canals lined by hymenium and filled at first by loose hyphae, these often breaking away raggedly in the wider cavities; peridium dark, minutely pseudoparenchymatous; gleba principally of compactly arranged hyphae or of prosenchyma. Hymenium consisting of asci and paraphyses in palisade, many of the paraphyses extending beyond the asci and filling the cavity with loose hyphae. Asci mainly cylindric or slightly clavate, normally 8-spored at first but with generally only 1-4 spores maturing; mature spores typically uniseriate, globose, smooth.
Desmazierella Lib.
Ann. Sci. Nat., sér. 1, 17: 83 (1829)
Type species: Desmazierella acicola Lib. 1829
Type designated by Libert (1829)
Diagnosis: Receptaculum orbiculatum; hymenium discoideum, discretum, setulis rigidis hirsutum ; Asci elongati, deorsum attenuati, flexuosi, absque paraphiysibus ; sporidia uniserialia, alba, ovata, sporidiolis duobus.
Family: Chorioactidaceae
Asexual morph of Verticicladium type.
Dichobotrys Hennebert
Persoonia, 7 (2): 193 (1973)
Type species: Dichobotrys abundans Hennebert 1973
Type designated by Hennebert (1973)
Diagnosis: Coloniae effusae, ochraceae hyphis hyalinis, laxis. Conidiophori erecti, magni, repetito dichotomice furcati; rami longi, divaricati, sursum curtiores, terminalibus geminatis globosis inflatis conidiogenis cellulis, simul conidia producentes, demum collabentibus. Conidia holoblastica, singula, sessilia vel pedunculata, subglobosa vel napiformia, laevia, superne saepe crassiori parite, inferne tenui pariete, cum vestigio pediculi dilabentia.
Synonym of Trichophaea
Dictyocoprotus J.C. Krug & R.S. Khan
Mycologia, 83 (1): 103 (1991)
Type species: Dictyocoprotus mexicanus J.C. Krug & R.S. Khan 1991
Type designated by Krug & Khan (1991)
Diagnosis: Apothecia dispersa, sessilia, levia, discoidea vel patelliformia, albida vel flavida. Excipulum iodo non rufescentes, ex excipulum ectale textura angulari et excipulum medullare textura intricata compositum, Asci protrudentes, operculati, iodo non caerulescentes, cylindracei, basin versus in stipitem brevem attenuati. Paraphyses filiformes, simplices, hyalinae, in apice crassae et uncinatae. Ascosporae unicellulares, ellipsoideae, hyalinae vel flavidae, subtiliter reticulatae, bullam unam continentes, sine vagina gelatinosa.
Family: Ascodesmidaceae
Current definition: Apothecla scattered, sessile, smooth, discoid to saucer-shaped, whitish to pale yellowish; excipulum cyanophilous, non-dextrinoid, two-layered, composed of an ectal excipulum of textura angularis and a distinct medullary excipulum of textura intricata. Asci protruding, operculate, nonamyloid, cylindrical, tapering below into a short stipe. Paraphyses filiform, simple, hyaline, enlarged and uncinate at the apices. Ascospores one-celled, ellipsoidal, hyaline to pale yellowish, ornamented with a fine reticulum, at maturity forming de Bary bubbles, lacking mucilaginous sheath.
Diehliomyces Gilkey
Mycologia, 46 (5): 789 (1954)
Type species: Pseudobalsamia microspora Diehl & E.B. Lamb. 1930
Type designated by Gilkey (1954)
Diagnosis: Ascomatibus hypogaeis vel epigaeis, a mycelio denso antecessis, e hypharum nodis efformatis, difformibus, connatis; ascis inter ascomatum hyphas stipatis, in plicis fertilibus a hyphis sterilibus separatis; sporis 8 vel paucioribus, haud regulariter ordinatis, ut pulvis luteus ab ascarum dissolutione liberatis.
Family: Otideaceae
Dingleya Trappe
Mycotaxon, 9 (1): 331 (1979)
Type species: Dingleya verrucosa Trappe 1979
Type designated by Trappe (1979)
Diagnosis: Ascocarpia exsiccata usque ad 3.5 cm in diam, subglobosa vel lobata. Peridium verrucosum, fragmentulis pubescentibus, brunneum. Gleba loculis labyrinthinis, hymeniis limitatis, sporis et ascis farctis. Asci cylindrici, pro parte maxima octospori, inamyloidei. Sporae ellipsoideae, 19-26 × 15-18 µm sine ornamentis; episporis 1-3 µm incrassatis, amorphis, brunneo-vinosis, nodosis et porcatis. Paraphyses ± 2 µm in diam, aequales vel inferiores quam asci. Holotypus: PDD 29712.
Family: Tuberaceae
Current definition: Ascocarps hypogeous with a brown, verrucose peridium. Gleba white with rounded to narrow, labyrinthine chambers lined with a hymenial palisade and filled with asci, paraphyses and spores. Asci cylindric, mostly 8-spored. Spores globose to ellipsoid, hyaline to brown, ornamented with cyanophilic, rounded to irregular warts or ridges.
Discina (Fr.) Fr.
Summa veg. Scand., section post.: 348 (1849)
Type species: Peziza perlata Fr. 1822
Type designated by Fries (1849)
Diagnosis: Cupula (ampla) ab initio plano-convexa, centro adfixa. Sporidia sporidiolis duobus.
Family: Discinaceae
Current definition: Ascomata epigeous, sessile to substipitate; hymenophore cupulate to discoid; hymenium rugose, yellow–brown, orange-brown, red-brown, or dark brown; stipe fluted, lacunose. Ascospores hyaline, ellipsoid to fusoid, with solitary, pointed apiculi; surface with an irregular reticulum at maturity; perispore cyanophilic; contents uniguttulate or triguttulate.
Discinella P. Karst., nom. illeg.
Hedwigia, 30: 301 (1891)
Type species: Discinella corticalis P. Karst. 1891
Type designated by Karsten (1891)
Diagnosis: Est Discina Fr. em. apotheciis minoribus. Complectitur species plures Humariae Sacc. Syll.
Family: Incertae sedis
Number of species: 1 fide DoF
Disciotis Boud.
Bull. Soc. mycol. Fr., 1: 100 (1885)
Type species: Peziza venosa Pers. 1801
Type designated by Eckblad (1968)
Diagnosis: Spores ovales elliptiques tardives, sans sporidioles. Genre fondé sur les P. venosa Pers. et reticulata Grev.
Family: Morchellaceae
Discomycetella Sanwal
Sydowia, 7: 200 (1953)
Type species: Discomycetella aquatica Sanwal 1953
Type designated by Sanwal (1953)
Diagnosis: Apothecia glabra, discoidea, hymenio convexo, valde gelatinoso. Hymenium in pagina superiore tantum evolutum. Asci longissimi, in stipitem attenuati, operculo apicali dehiscentes. Sporae hyalinae, ellipsoideae leves. Mycelium vegetativum superficiale, niveum
Synonym of Psilopezia?
Dissingia K. Hansen, X.H. Wang & T. Schumach.
Persoonia, 42: 195 (2019)
Type species: Peziza leucomelaena Pers. 1822
Type designated by Hansen et al. (2019)
Family: Helvellaceae
Current definition: Apothecia cup-shaped, mostly remaining concave when expanding, subsessile, or with a ± distinct stipe; stipe short, broad or slender, with a few ± conspicuous, blunt ribs gradually widening and subdividing towards the cup attachment. Hymenium yellowish brown to greyish brown, to dark brownish black; receptacle surface even or with few to many blunt to angular ribs and grooves below, pubescent, upper part whitish to dark greyish brown, below concolorous or often gradually paler to almost whitish. Asci cylindrical, operculate, arising from simple septa, 8-spored. Spores ellipsoid, obtuse or attenuate, smooth. Paraphyses filiform, septate, with clavate or subcapitate tips.
Donadinia Bellem. & Mel.-Howell
Cryptog. Mycol., 11 (3): 218 (1990)
Type species: Urnula helvelloides Donadini, Berthet & Astier 1973
Type designated by Bellemère et al. (1990)
Diagnosis: Apotheciae lignicolae, in ligno mortuo caduco enascentes, pedunculatae, cupulares, gregariae; cupula hemisphaerica, marginata. Stipes plenus. Asci octospori, cylindrati, base flexuosi. Endotunica in apice solum adest, natura heteroclita. Ectotunica in apice angustata, natura heteroclita. Apertio operculata haud suboperculata. Sporae pariete propra dispari, ac perispora verrucis parvis cyanophilisque ornata, et intus parcis guttulis repletae. Ascogenae hyphae non fibulatae. Paraphyses lineares, apice saepe furcatae ac nodosae.
Family: Sarcosomataceae
Durandiomyces Seaver
N. Amer. Cup-Fungi (Operc.): 242 (1928)
Type species: Gyromitra phillipsii Massee 1895
Type designated by Seaver (1928)
Diagnosis: Ascophores large, consisting of numerous contorted and frequently anastomosing ribs resembling a Spamssis, the numerous cavities lined everywhere with the hymenium, the color ranging from white to rosy or faintly violaceous; flesh very brittle; asci 8-spored; spores minutely sculptured; spore-sculpturing consisting of minute warts or papillae.
Synonym of Daleomyces
Elaiopezia Van Vooren
Ascomycete.org, 12 (4): 185 (2020)
Type species: Galactinia polaripapulata J. Moravec 1969
Type designated by Van Vooren (2020)
Family: Pezizaceae
Current definition: Ascomata epigeous, discoid or slightly cupulate, small-sized, olive or yellowish with distinct olivaceous tinges. Flesh without latex. Asci operculate, 8-spored; ascus wall diffusely amyloid (W type). Paraphyses containing a yellow, yellow-ochre or olivaceous pigment, sometimes also with an external pigment at the top. Ectal excipulum of textura globulosa with small to medium-sized cells. Ascospores eguttulate, sometimes containing small granules or drops in young state, smooth or ornamented. Species saprobic.
Eleutherascus Arx
Persoonia, 6 (3): 377 (1971)
Type species: Arachniotus lectardii Nicot 1970
Type designated by Arx (1971)
Diagnosis: Ascomata absunt; ascogonia in mycelio aerio, curvata vel contorta, ascos singulos vel paucos proferunt et saepe nonnulla filamenta; asci late clavati, obovati vel subglobosi, pariete simplici, tenui, plerumque octospori; ascosporae globosae, maturitae dilute brunneae, pariete crasso, spinoso; conidia absunt.
Family: Ascodesmidaceae
Eoaleurina Korf & W.Y. Zhuang
Mycotaxon, 26: 381 (1986)
Type species: Humaria foliicola Henn. 1902
Type designated by Zhuang & Korf (1986)
Diagnosis: Ab Aleurina Massee cellulis excipuli ectalis isodiametricis, globosis vel angularibus, et non polygonaliter elongatis, parietibus cellularum exilibus et hyalinis; non rigidis et pigmentiferis differens. Ab Greletia Donadini et Smardaea Svrcek contentu cellularum excipularum brunneo, non purpureo, in speciminibus siccatis in aqua ad examinationem paratis, et apicibus paraphysium tumidis differens. Ab his generibus trbus omnibus texturis apothecialibus parum gelatinosis, et excipulo medulloso ex permixtione texturae angularis et texturae intricatae potius quam tantum ex textura intricata non gelatina consistente differens.
Family: Pyronemataceae
Number of species: 1 fide DoF
Current definition: Apothectum dlscoid to subturbinate, sessile to subsessile, hymenium smooth, receptacle pustulate. Excipular tissues slightly gelatinous; ectal excipulum of textura globulosa to textura angularis, cells hyaline, thin-walled, pigmentation if present only in the cytoplasm; medullary excipulum of textura angularis mixed with textura 1ntricata. Subhymenium indistinguishable. Asci operculate, 8-spored, subcylindrical, J- in Melzer''s Reagent. Ascospores ellipsoid, biguttulate, with ornamentations on the surface. Paraphyses swollen at apex, with pigmented contents.
Eoterfezia G.F. Atk.
Bot. Gaz., 34: 40 (1902)
Type species: Eoterfezia parasitica G.F. Atk. 1902
Type designated by Atkinson (1902)
Diagnosis: Fruit bodies subglobose, minute, fleshy, with a distinct but very thin, undifferentiated wall, smooth or slightly hairy. Interior of fruit body with sterile avenues radiating from the base and branching until they unite with the wall. Fertile areas lying between containing the asci intermingled with the supporting hyphae. Spores smooth, hyaline, one-celled. Interior not disintegrating into a powdery mass at maturity. At present one species as follows: […]
Family: Incertae sedis
Number of species: 2 fide DoF
Eremiomyces Trappe & Kagan-Zur
Mycol. Res., 109 (2): 244 (2005)
Type species: Choiromyces echinulatus Trappe & Marasas 1973
Type designated by Ferdman et al. (2005)
Diagnosis: A Choiromyces sporis virgis conicis rectis obtusis, cellulis magnopere inflatis, ordiniibus DNA abhorrentibus (ad Pezizaceas, non Tuberaceas, affinis) et distributione in desertis Hemisphaerio Meridionali differt.
Family: Pezizaceae
Fallacidiscus U. Lindemann, Roffler & Van Vooren
Ascomycete.org, 14 (2): 46 (2022)
Type species: Fallacidiscus helveticus U. Lindemann, Roffler & Van Vooren 2022
Type designated by Lindemann et al. (2022)
Diagnosis: Differs from Spaniodiscus fuckelii and Ramsbottomia spp. by discoid apothecia with a small marginal edge, a hairless margin, and finely warted ascospores containing polar oil droplets, as well as its genetic profile.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, yellow-orange-coloured, with an external surface densely covered by short brown hairs and a hairless margin. Excipulum two-layered: medullary layer of textura intricata, ectal layer of textura subglobulosa/angularis. Hairs short, simple, superficial, septate, with a simple base, obtuse or slightly sharp. Asci operculate, 8-spored, narrowing toward base, arising from croziers, inamyloid. Paraphyses slender, ± enlarged at the top, containing carotenoid pigments. Ascospores uniseriate, ellipsoid, hyaline, thick-walled, containing small bipolar oil droplets, finely warted. Trophic status unknown. Asexual morph unknown.
Fastigiella Benedix
Kulturpflanze, 17: 276 (1969)
Type species: Morchella caroliniana Bosc 1811
Synonym of Neogyromitra
Filicupula Y.J. Yao & Spooner
Kew Bull., 51 (1): 193 (1996)
Type species: Pseudonectria suboperculata Döbbeler & P. James 1978
Diagnosis: Apothecia primum clavata, demum obovoidea vel turbinata, supra applanata. Discus aurantiacus. Receptaculum cupulatum, ad basin hyphis ramosis affixum. Excipulum ectale ad basin e textura prismatica compositum, ad lateris e hyphis subparallelis septatis compositum, muris hypharum crassis gelatinosisque; hyphae ad angulum parvum ad superficiem depositi. Asci operculati, jodo non caerulescenti, clavati. Ascosporae unicellulares, hyalinae, laeves, fusoideae, guttulatae. Paraphyses filiformes, septatae, rectae vel curvatae.
Family: Incertae sedis
Current definition: Apothecia clavate at first, becoming obovoid or turbinate, flattened above. Disc orange. Receptacle cupulate, surface rough, with branched anchoring hyphae present at the base and lower receptacle. Ectal excipulum comprising textura prismatica at the base, and septate, sub-parallel hyphae above; excipular hyphae radially arranged, lying at a low angle to the surface, their walls thickened and strongly adherent. Asci operculate, iodine-negative, clavate, thin-walled. Ascospores unicellular, colourless, smooth, fusoid, often inaequilateral, guttulate. Paraphyses filiform, septate, straight or curved, flexuous.
Fischerula Mattir.
Nuovo G. bot. ital., 34: 1348 (1928)
Type species: Fischerula macrospora Mattir. 1928
Type designated by Mattirolo (1928)
Diagnosis: A Tuberaceis adhuc notis eximium hoc novum Genus, ascorum, sporarumque habitu differt, dum facies universa cum aliis fungis hypogeais (praesertim e Genere « Tuber ») plerumque convenit. Peridio irregulari, difformi, circumvoluto, contextu pseudoparenchymatico ; colore umbrino-luteo notato. Carne fuliginea, venis luteis filiformibus dispersis. Ascis amplis, membranis spissis, formam praebentibus ovatam (130 usque ad 179 micra et ultra long, secundum sporarum inclusarum numerum) basi leviter attenuatis. Sporis raro unicis; plerumque 2-3-4 in singulis ascis, magnis (50-70 long. et ultra ; 40-50 lat. et ultra); non aculeatis, nec alveolatis (uti in Genere « Tuber »); sed dense verrucosis; verrucis difformibus, irregularibus, latis ; colore saturate castaneo-badio, illum Hydnothriae quodammodo referente.
Family: Morchellaceae
Fuckelina Kuntze, nom. illeg.
Rev. Gener. Plant., 2: 852 (1891)
Type species: Octospora bulbosa Hedw. 1788
Diagnosis: = Macropodia Fuck. 1869 non (—um) R.Br* 1812 = Octospora Hedw. ex parte minima. Die Crucifere Macropodium gilt, sodass die homonyme Discomycetee anders zu benennen ist; sie sei dem Begründer dieser Gattung gewidmet. Die Arten sind nach Saccardo syll. VIII 158—160 von Macropodia übertragen [...]
Synonym of Helvella
Galactinia (Cooke) Boud.
Bull. Soc. mycol. Fr., 1: 101 (1885)
Type species: Peziza saniosa Schrad. 1792
Type designated by Clements & Shear (1931)
Diagnosis: Cupula sessilis, integra, lactescens
Family: Pezizaceae
Galeoscypha Svrček & J. Moravec
Česká Mykol., 43 (4): 210 (1989)
Type species: Coprobia pileiformis Svrček 1978
Type designated by Svrček & Moravec (1989)
Diagnosis: Carposomata parva, gregaria, usque confluentia, superficialia, absque hypothallo sed basi hyphis ferrugineo-brunneis, tenuibus, crasse tunicatis, septatis, dense connexis substrato adnata, primum subglobosa, pyriformia, apice truncata, dein pileiformia, disco convexo, hemisphaerico, immarginato, nonnumquam obtuse conico, margine undulato, granuloso, fulgide aurantiaco, subtus crasse obconica vel subcylindrica, stipitiformiter angustata, thecio sanguineo-rubro vel aurantiaco-rubro, extus minutissime granulata Excipulum externum atque medullaris textura globuloso-angularis, aurantiaco-coloratum, parte externa margineque cellulis globosis cyanophilis, ad basim pilis hyphoideis flexuosis hyalinis septatis et hyphis mycelialibus ferrugineo-brunneis instructum. Asci cylindrici, operculati, non amyloidei, octospori. Paraphyses crasse cylindricae, parte basali ramosae, sursum dilatatae, clavatae, subcapitatae vel lanceolatae, septatae, sub hyalinae. Ascosporae ellipsoideae, inaequales, intus irregulariter granulosae sed etiam biguttulatae, perisporio non separabile, verrucis cyanophilis in aequalibus ornato. Habitat saprobioticus.
Family: Pyronemataceae
Galiella Nannf. & Korf
Mycologia, 49 (1): 107 (1957)
Type species: Bulgaria rufa Schwein. 1832
Type designated by Korf (1957)
Diagnosis: Apothecia epixyla, plerumque magna, sessilia vel breviter stipitata, carne egregie crassa gelatinosaque, plerumque hemisphaerica vel crateriformia; sicca maxime contracta, facie plicata vel rugosa. In sectione hypothecium plerumque distinctum; excipulum medullare e textura intricata in matrice gelatinosa sita, saepe crassissimum, aliquando hypharum parallelarum zonis praeditum ; excipulum exterum e textura intricata vel ad texturam angularem vel globulosam vergens, cellulis brunneo-tunicatis, capillas gignens. Capillae brunneo-tunicatae, longae vel breves, leves vel asperae, flexuosae, specie viventes. Hymenium color pallido ad rufum vel griseo-atrum vergente, aliquando, praecipue siccum, fere atrum. Asci magni, longi, deorsum attenuati, modice crasse tunicati, suboperculati, liquore iodi haud cyanescentes. Ascosporae uniseriatae, unicellulares, magnae, punctis e materie callosa-pectica tenuibus vel crassis ornatae. Paraphyses graciles, interdum ramosae vel anastomosantes.
Family: Sarcosomataceae
Genabea Tul. & C. Tul.
G. bot. ital., 1 (7-8): 60 (1845)
Type species: Genabea fragilis Tul. & C. Tul. 1845
Type designated by Tulasne & Tulasne (1845)
Diagnosis: Peridium distinctum nullum. Moles carnosa firma irregularis globosa gibberoso-tuberculosa sinuosa, quandoque hinc et illinc anguste rimosa vel pertusa, absque basi radicali, intus cuniculis gyrosis cavis, ad rimas vel aperturas externas, quarum opera aerem admittunt, pertingentibus, parce exsculpta percursa; contextu celluloso densissimo. Sporangia composita concreta et aggregata, sunt nempe nil nisi loculamenta oblonga septis tenuibus indivisis obscuris distincta, norma quadam conjuncta, quae spatia tenent propria glomerulosque efficiunt nunc globosos nune cylindricos aut reniformes huc et illuc in mole remote nidulantes nec ab ipsa solubiles, reliqua substantia sterili avenia dilutius colorata. Sporae crassae ovatae glabrae leves nitentes et quasi crustaceae, 4-6 vel pauciores in quocumque loculamento.
Genus Hydnoboliti et Geneae quodam modo affine.
Family: Pyronemataceae
Genea Vittad.
Monogr. Tuberac.: 27 (1831)
Type species: Genea verrucosa Vittad. 1831
Diagnosis: CHAR. Uterus rotundatus, difformis, tuberoso-plicatus, subcavus, apice pervius; peridium crassum, extus intusque flоссоsum vel papilloso-muricatum, hinc illinc duplicato-intrusum. Sporangia cylindracea, octospora ! in peridii parte media carnosa, transversim ac parallele disposita. Sporidia albida subrotunda.
» Obs. Uterus vix subterraneus, e peridio parum vel laxe intruso subcavus, fibrillis radicalibus corymbosis instructus, vel floccis undique cinctus, apice rima, margine spisso circumdata, in cavo uteri adaperta omatus. Sporidia opaca subechinata, seu granulis veluti adspersa.
HIST. Fungi minores, graveolentes, elengantissimi, persistentes, Tuberibus, structura quamvis simpliciori ac veluti prototypa, affines; sporangiorum vero dispositione et natura distincti. Pezizis forma, tum etiam fructificatione, etsi vere uterina, analogi.
Family: Pyronemataceae
Geneosperma Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 102 (1968)
Type species: Geneosperma geneosporum Rifai 1968
Type designated by Rifai (1968)
Diagnosis: Apothecia gregaria, mediocria, rubro-aurantiaca, sessilia, patelliformia, hirsuta. Pili rubro-brunnei, crasse tunicati, septati, acuminati, basi saepe appendice radiciformi aucti. Excipulum ectale ex cellulis crassis angularibus (textura angulari), excipulum medullare ex hyphis septatis compositum. Asci subcylindracei, octospori, apice jodo haud tincti. Ascospori uniseriati, hyalini, ellipsoidei, guttulati, verrucosi, utrinque appendiculati, processibus utrinque acutis lateraliter subappressis et apicibus conoideo-folliculatis. Paraphyses septatae, superne rectae, clavatae.
Hab. ad lignum putridum inter muscos in India orientali, Malesia. et Japonia.
Family: Pyronemataceae
Number of species: 1 fide DoF
Geodina Denison
Mycologia, 57 (4): 649 (1965)
Type species: Geodina guanacastensis Denison 1965
Type designated by Denison (1965)
Diagnosis: Apothecia cupularia, stipitata; stipes angustum; excipulum crassum, subereum vel coriaceum; intus filamentosum, extra membranaceum e cellulis subrectangularibus constans; hymenium pallidum vel coloratum, luteum vel rubrum; extra simile colore et pilosum; pili fasciculati ex hyphis crasse tunicatis et continuis; asci suboperculati, cylindracei, in basim sensim angustati, iodo addito non caerulei, non simul maturescentes; opercula eccentrici; sporae ellipsoideae, amplae, 1-2 guttis adfectae, costis magnopere sculptae; paraphyses tenues cylindraceaeque, septatae, in apice clavatae. Plantae terrestres, clima calido vigentes.
Family: Wynneaceae
Current definition: Apothecium goblet-shaped with a slender stem, thick-fleshed, corky to leathery; hymenium light-colored to bright-colored, yellow to red; exterior similarly colored, hairy; hairs fasciculate composed of thick-walled, nonseptate hyphae arising from the ectal excipulum; excipulum composed of two distinct layers, a medullary excipulum of entangled hyphae (textura intricata), and a rind-like ectal excipulum composed of more or less rectangular cells (textura prismatica to textura angularis); asci suboperculate, with eccentric opercula, cylindrical with gradually tapering bases, not turning blue in iodine, not all maturiilg simultaneously; ascospores ellipsoid, large, long, containing 1-2 large oil drops, heavily sculptured; spore sculpturing not staining readily in cotton blue, consisting of coarse, anastomosing ridges; paraphyses slender, cylindrical, septate, with subclavate apices; growing in the soil in the tropics.
Geomorium Speg.
Anal. Soc. cient. argent., 94 (1-2): 79 (1922)
Type species: Geomorium fuegianum Speg. 1922
Type designated by Spegazzini (1922)
Diagnosis: Char. Helvelleum; stipes carnoso-tenacellus albescens, sursum saepius subattenuato-rotundatus, late fistulosus, cavitate interna septis longitudinalibus plus minusve anastomosantibus corroboratus, extus verticaliter subirregulariter costulatus, in parte dimidia supera hymenio effuso arcte adnato atro-olivaceo vestitus; asci cylindracei, apice rotundati (porosi?) octospori. paraphysibus bacillaribus tenuibus sursum vix inerassatis simplicibus intus olivaceo-granulosis obvallati; sporae ellipticae, hyalinae, verrucosae.
Genus Morchellam accedens, a qua ascomatis hymeniique fabrica, trabeculis transversis deficientia, nec non sporis verrucosis recedit.
Family: Geomoriaceae
Current definition: Ascomata either a modified columnar apothecium or an exothecium, 1–40 mm broad. Young ascomata typically white or pale tan but changing to brown, purple, or black as the hymenium matures. In the epigeous species, lacking hairs, outer excipulum a palisade-like layer, 50–350 μm thick, made up with isodiametric cells, 7–20 μm diam, perpendicular to the outer surface. Paraphyses typically slightly exceeding the length of asci, although paraphyses far exceed the asci in some hypogeous taxa. At maturity in some taxa paraphyses have dark granular inclusions in the apical cells. Asci cylindrical, spores uniseriate in epigeous taxa, but often irregularly uniseriate or biseriate in hypogeous taxa, lacking opercula. Ascospores typically hyaline but occasionally yellowish, ellipsoid to subglobose, and ornamented with warts that range from small in the epigeous taxa to large in the hypogeous taxa. Ectomycorrhizal, found in forests or at forest edges, fruiting on the ground in association with Nothofagaceae (South America and Australasia) or Myrtaceae (Australasia) and perhaps with other host plants. Known only from the Southern Hemisphere.
Geopora Harkn.
Bull. Calif. Acad. Sci., 1 (3): 168 (1885)
Type species: Geopora cooperi Harkn. 1885
Type designated by Harkness (1885)
Diagnosis: Subterranean. Integument woolly, continuous with the trama. Hymenium convolute. Asci cylindrical. Sporidia hyaline, oblong, smooth.
Family: Pyronemataceae
Geoporella Soehner
Z. Pilzk., 21 (8): 8 (1951)
Type species: Geopora michaelis E. Fisch. 1898
Diagnosis: Ascomata subglobosa, tuberosa, lobata; cortice plerumque in glebam profunde penetrante; superficie ab initio cerea, mature fusco-purpurea; gleba purpurea, cavernis trajekta et gyrum carpta; asci cylindrici, oktospori; asci subhymeniales copiose prominentes; sporae uniseriatae, ellipsoideae, verrucosae vel echinatae papillosae; paraphyses ascos superantes.
Synonym of Hydnotrya
Geopyxis (Pers.) Sacc.
Syll. fung. (Abellini), 8: 63 (1889)
Type species: Peziza carbonaria Alb. & Schwein. 1805
Type designated by Korf (1972)
Diagnosis: Cupula integra, sessilis aut stipitata stipite laevi
Family: Tarzettaceae
Current definition: Apothecia discoid, cupulate to urnulate, 3–25 mm in diam, 2–18 mm high, fleshy, sessile to stipitate, stipe up to 14 mm long 6 3 mm thick. Hymenium yellow, orange, orange-red, saffron, ochraceous buff, brownish yellow; receptacle surface concolorous with hymenium or darker, smooth or with brown warts; margin raised, crenulate, cream-colored to white. Asci cylindrical, operculate, 8-spored, hyaline, base with croziers. Spores uniseriate, ellipsoid to subfusiform, hyaline, smooth, sometimes ornamented with very fine warts or low ridges, eguttulate. Paraphyses of equal width or slightly enlarged at apices, mostly straight, when fresh with yellow to orange-red granules or guttules, when dried with dense or sparse hyaline granules. Ectal excipulum of textura angularis; cells thick-walled, hyaline or with yellowish-brown walls. Medullary excipulum of textura intricata, hyphae hyaline or with amber-like resinous exudates, not dissolving or changing in KOH or MLZ.
Substrate: On burned or unburned ground.
Geoscypha (Cooke) Lambotte
Mém. Soc. roy. sci. Liège, 2e sér., 14: 320 (1888)
Type species: Peziza violacea Pers. 1794
Type designated by Eckblad (1968)
Diagnosis: Cupula integra, vel lobata, sessilis, extus laevi, sub-pruinosa vel farinosa
Family: Pezizaceae
Current definition: Ascomata epigeous, cupulate or discoid, violet-coloured. Asci operculate, 8-spored, with the top intensely bluing in iodine solution (WTR or WT type in G. ampelina), with crozier. Paraphyses containing a brown or purple brown vacuolar pigment. Ectal excipulum of textura globulosa, with large cells. Ascospores smooth or ornamented with isolated warts, biguttulate, except in G. violacea where the drops may be visible only in young spores. Species saprobic, often growing on burnt soil.
Gilkeya M.E. Sm., Trappe & Rizzo
Mycologia, 98 (5): 705 (2007)
Type species: Hydnocystis compacta Harkn. 1899
Type designated by Smith et al. (2007)
Diagnosis: A Genea sporis subglobosis globosis, peridio vinaceo et absentia caespitis basalis hypharum differt.
Family: Pyronemataceae
Glaziella Berk.
Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn, 80: 31 (1880)
Type species: Glaziella vesiculosa Berk. 1880
Type designated by Berkeley, in Glaziou (1880)
Diagnosis: Stroma subglobosum laeticolor; perithecia pallida, gelatina hyalina repleta.
Family: Glaziellaceae
Globopilea Beauseign.
Contr. Etude Fl. Mycol. Landes: 204 (1926)
Type species: Helvella phlebophora Pat. & Doass., in Patouillard 1886
Type designated by Beauseigneur (1926)
Synonym of Helvella
Greletia Donadini
Bull. Soc. mycol. Fr., 95(3): 183 (1980)
Type species: Plicaria planchonis Dunal ex Boud. 1887
Type designated by Donadini (1980)
Diagnosis: Apothecia 0,5-4 cm, substipitata vel subsessilia, cupulata, terricola vel humicola. Hymenium semper caesium vel violaceum. Excipulum concoloratum. Caro violacea. Pigmentum violaceum in omnibus fungi partibus (asci, sporae, paraphyses, caro, mycelium). Asci jodo non caerulescentes. Excipulum medullosum textura intricata compositum et ectale excipulum textura globulosa angularis constitutum. Sporae in cumulo albae saepe albolilacinae.
Family: Pyronemataceae
Number of species: 5
Gymnohydnotrya B.C. Zhang & Minter
Mycol. Res., 92 (2): 192 (1989)
Type species: Gymnohydnotrya australiana B.C. Zhang & Minter 1989
Type designated by Zhang & Minter (1989)
Diagnosis: Ascomata subterranea, subglobosa vel irregularia, 15-25 mm diam vel dimensionis maximae respectu, lobata, parvis et conpluribus cavitatibus et canaliculis quae in superficiem sese aperiuntur, vel una cavitate induta, laevia et in fundamento quaeque cum flocculo. Peridium absens. Hymenium exteriorem totam et saepius interiorem tegens. Paraphyses cylindricae, saepius dichotome ramosae, septatae, apicibus cum liberis, rotundatis et inflatis, nil vel ascos vix superantes. Asci cylindrici, nil in iodo caerulescentes, nec dehiscentes, plurima parte octospori, sed nonnumquam cum una vel duabus ascosporis abortis. Ascosporae uniseriatae, ellipsoideae, sine colore, quadrinucleatae, cum reticulis instar fornicibus vel spinis et pariete cyanophilicis.
Synonym of Geomorium
Gyrocephalus Pers., nom. rejic.
Mém. Soc. Linn. Paris, 3: 77 (1824)
Type species: Gyrocephalus aginnensis Pers. 1824
Diagnosis: Pileus ? (aut capitulum) tremellosus aut subtremellosus, gyroso-sinuosus, suffultus stipite forti.
Synonym of Gyromitra
Gyrocratera Henn.
Verh. bot. Ver. Prov. Brandenb., 41: 9 (1899)
Type species: Gyrocratera ploettneriana Henn. 1899
Type designated by Hennings (1899)
Diagnosis: Tuberacearum. Ascomata carnosa, rotundata, glabra vel gyroso-lobata, laevia, apice foramine rotundato, arrhiza; gleba nune lacuna amplissima simplici excavata, nunc varie cavernosa, anfractibus labyrintheis continuis in ostium apicale simul confluentibus. Asci cylindraceo-clavati, 6—8 sporis; paraphysibus paulo superantibus, apice liberis. Sporae crassae verrucosae, brunneae, subglobosae.
Synonym of Hydnotrya
Gyromitra Fr.
Summa veg. Scand., section post.: 346 (1849)
Type species: Helvella esculenta Pers. 1800
Type designated by Clements & Shear (1931)
Diagnosis: Helvellae sp. Auctt. Discus bullato-inflatus, costis elevatis gyrosus.
Family: Discinaceae
Current definition: Ascomata epigeous, stipitate; hymenophore irregularly convex to irregularly lobed or cerebriform; hymenium rugose to convoluted, orange-brown to dark red-brown to purplish; stipe cylindric, sometimes with longitudinal furrows, hollow, white to cream, brown or purple. Ascospores hyaline, ellipsoid to subfusoid, nonapiculate or with broadly rounded apiculi; surface seen as smooth with light microscopy but finely rugose under SEM at maturity; perispore cyanophilic; contents biguttulate.
Gyromitrodes Vassilkov
Sovetsk. Bot., 6: 51 (1942)
Type species: Helvella sphaerospora Peck 1875
Synonym of Pseudorhizina
Hansenopezia Matočec, I. Kušan & Jadan
in Yuan et al., Fungal Divers., 104 : 64 (2020)
Type species: Peziza retrocurvata K. Hansen & Sandal 1998
Type designated by Matočec et al. (in Yuan et al., 2020)
Family: Pezizaceae
Current definition: Ascomata apothecial, solitary or often in groups, medium-sized, epigeous, centrally attached to the substrate, regularly circular to much sinuous from the top view, cupulate when young, then plate-shaped, hymenial surface reddish brown, lilac-violet, violet-brown, purplish brown to dark brown, matte, margin upright then flattened to reflexed, entire, blunt, smooth and or beset with scattered white fibrils, whitish grey or pale brownish yellow, excipular ground concolourous with the margin, finely pubescent or ornamented with reddish brown patches distributed to the basal parts. Flesh fragile, homogenous, soft waxy, dry on cut or exuding watery to milk-white latex. Hymenium arranged as a regular palisade. Asci cylindrical, hyaline, apex subtruncate, protruding above paraphysis tips at full maturity, 8-spored, base tapered, arising from croziers, operculum apical, very narrow, occupying ≤ 28% of living mature ascus diam, functional, flat lentiform before opening, encircled by slight “u”-shaped indentation in the wall, periascal amyloid mucus thin and film-like through the entire length but almost lacking on opercular lense, while the thickest at the opercular edge where is moderately euamyloid, forming unclear thin annular zone, reaction gradually decreasing in strength towards the base, with reacting area on half of the ascus length or more. Ascospores hyaline, ellipsoid to narrowly ellipsoid with ± rounded poles, radially symmetrical, 1-celled, subglabrous or with cyanophilic verrucae, perispore cyanophilic, not loosening when heated in lactic Cotton Blue or in KOH, wall 3-layered ± thick-walled, without sheath on ejected spores; sporoplasm multiguttulate, lipid content nearly maximal with all guttules of approx. same size or 2–3 large lipid bodies mixed with many smaller ones, lipid bodies much coalesced in dead spores, uninucleate, nucleus and nucleolus not contrasted nor stained in Lugol’s solution, de Bary bubbles absent in anhydrous media. Paraphyses cylindrical, apically obtuse to subclavate, straight to bent, not branching in the upper part, embedded in hyaline epithecial exudate or only with hyaline, thin coating, apical cell contain single or few non-refractive vacuoles entirely filled with finely dispersed minute pinkish brown particles and/or partially with larger non-refractive granules of the same colour, not containing vacuolar bodies, in Lugol’s solution pigment partially turning to rusty brown colour. Marginal texture in section displaying broad outline, textura globulosa-angularis mixed with some cylindric cell chains that may protrude as scattered cylindric obtuse, thin-walled hyphoid hairs or form loose tufts; texture is not cyanophilous. Subhymenium well-discerned, composed of hyaline subgelatinous densely woven textura porrecta. Medullary and ectal excipulum of ± homogenous, subhyaline texture, except for the cortical layer. Upper part composed of textura globulosa- moniliformis mixed with cylindric very long, mostly vertically oriented hyphae. Lower part composed of textura globulosa-angularis with intermixed intricate connective hyphae, cells smaller in size towards the surface. Cortical layer composed of textura porrecta-intricata, discontinuous layer when present pustulate, cells with ochraceous walls and rusty brown patches, coiled and agglutinated together; texture is not cyanophilous.
Ecology: Saprotroph on rich soil with litter and wood residues of Abies alba, on very rotten stumps and trunks of Picea abies and Fagus sylvatica, sometimes also on rich soil outside forests, fruitbodies appear in spring or autumn to winter.
Hapsidomyces J.C. Krug & Jeng
Mycologia, 76 (4): 748 (1984)
Type species: Hapsidomyces venezuelensis J.C. Krug & Jeng 1984
Type designated by Krug & Jeng (1984)
Diagnosis: Apothecia dispersa, cleistohymenialia, turbinata, margine distincto, brunnea, excipulum ectale et excipulum medullare exhibentes. Asci protrudentes, operculo disciformi, iodo caerulescentes, cylindracei, flavidiores, basin versus in stipitem brevem contracti. Paraphyses filiformes, granula flavida continentes. Ascosporae unicellulares, globosae, eguttulatae, hyalinae vel pallide flavidae, reticulum et spinae exhibentes.
Family: Pezizaceae
Hellenicoscyphus U. Lindemann, Van Vooren & Kaounas
Ascomycete.org, 14 (2): 74 (2022)
Type species: Hellenicoscyphus hyalotrichus U. Lindemann, Van Vooren & Kaounas 2022
Diagnosis: Differs from Paratricharina by its hyaline marginal hairs organised in small bundles, the lack of excipular hairs and from Tricharina by its hyaline marginal hairs, the lack of excipular hairs and its bipolar spore granules, and from both genera by its genetic profile.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, cupuliform, light brown to ochre. Receptacle hairless. Marginal hairs organised in small bundles, superficial, septate, with a simple widened base. Excipulum two-layered: medullary layer of textura intricata, and ectal layer of textura globulosa/angularis. Asci operculate, narrowing toward base, arising from free croziers, inamyloid, 8-spored. Paraphyses slender, hyaline. Ascospores uniseriate, ellipsoid, hyaline, smooth, containing small bipolar granules. Species probably saprobic. Asexual morph unknown.
Helvella L.
Sp. pl., 2: 1180 (1753)
Type species: Helvella mitra L. 1753
Type designated by Dissing (1966)
Diagnosis: ELVELA pileo deflexo adnato lobato difformi. Fl. suec. 1103.
Fungoides fungiforme crispum laciniatum & varie complicatum, Mich. gen.-204. t. 96. f. 7.
Fungus autumnalis velut apex flaminis. Mentz. pug. t. 6.
Habitat in Truncis putridis.
Family: Helvellaceae
Current definition: Ascocarps (apothecia) in Helvella are either cupulate or capitate, subsessile or usually distinctly stipitate; when cupulate, cup deeply cupulate to saucer-shaped to discoid (planar), sometimes cup laterally compressed; when capitate, cap regularly campanulate to bi- to trilobate to irregularly saddle-shaped with cap edge free or partly attached to stipe, apothecial margin (cup margin or cap edge) recurved (deflexed) or adnate (reflexed) towards the stipe and sometimes firmly fused with it; hymenium whitish to greyish to brownish to black when fresh, receptacle surface (apothecial underside) concolorous or discoloured, glabrous or pubescent; stipe when present terete or ribbed and furrowed, inside solid or hollow or chambered. In some cupulate species the ribbed stipe becomes much reduced so that almost sessile apothecia occur. Asci are cylindrical, 8-spored, operculate, with an aporhynchous or pleurorhynchous base. Ascospores ellipsoid to ellipso-fusoid, hyaline, generally unsculptured or minutely verrucose in fully mature ascospores, containing a large central guttule when mature, tetranucleate. Paraphyses filiform, septate, not much inflated, or inflated (enlarged) at tips to a clavate or subcapitate apex.
Helvellella S. Imai
Bot. Mag. (Tokyo), 46: 174 (1932)
Type species: Helvella sphaerospora Peck 1875
Type designated by Imai (1932)
Diagnosis: Genus affine Helvellae sed sporae globosae
Synonym of Pseudorhizina
Heteroplegma Clem.
Bull. Torrey bot. Club, 30: 92 (1903)
Type species: Heteroplegma caeruleum Clem. 1903
Type designated by Clements & Shear (1931)
Diagnosis: Apothecia superficialia, sessilia, carnosa, hemisphaerico-cupulata, furfuraceo-excipulata, magna; epithecium nullum; paraphyses praesentes; hypothecium percrassum, hyalinum, trilaminatum, lamina superiore et inferiore pseudoparenchymaticis, medullari hyphis parallelis intertextisque, tramitiforme; excipulum tenue, filis brevibus tomentosum. Asci octospori, cylindracei, iodo valde caerulescentes. Sporae continuae, hyalinae, ellipticae. Nomen propter hypothecium laminatum, ἕτερος, dissimilis, πλέγµα, τό, vimineum.
Est Piicaria hypothecio heteromorpho.
Synonym of Galactinia
Heydenia Fresen.
Beitr. Mykol., 2: 47 (1852)
Type species: Heydenia alpina Fresen. 1852
Type designated by Fresenius (1852)
Diagnosis: Stipes erectus eximie cellulosas, e cellulis parenchymaticis et elongatis compositus, apice extus in marginem disciformem expansus, intus in columellam elevatus, capitulo floccorum septatorum sporigerorum subgloboso ornatus. Sporae simplices concatenato-conglomeratae isthmis brevissimis conjunctae, pleurogenae.
Family: Pseudombrophilaceae
Hiemsia Svrček
Česká Mykol., 23: 83 (1969)
Type species: Lachnea pseudoampezzana Svrček 1948
Type designated by Svrček 1969
Diagnosis: Apothecia solitaria usque gregaria, minuta, discoidea, late sessilia, disco plano, rubro-aurantiaco, margine subtiliter fimbriata, extus nigrofusco-fibrillosa. Excipulum hyphis umbrino-fuscis, longe cylindraceis, flexuosis, septatis, apice obtusis vel clavatis, constat. Medulla e hyphis angustioribus hyalinis composita. Asci cylindracei, crasse longeque stipitati, octospori, membrana non amyloidea instructi. Paraphyses crassae, septatae, apice subclavatae, rectae. Sporae obtuse ellipsoideae, guttula unica magna instructae, minute verrucosae, hyalinae.
Hab. Inter muscos humiles ad lapides.
Synonym of Octospora
Hoffmannoscypha Stielow, Göker & Klenk
Mycol. Progr., 12 (4): 684 (2012)
Type species: Scutellinia pellita Cooke & Peck ex Kuntze 1891
Type designated by Stielow et al. (2012)
Family: Pyronemataceae
Current definition: Apothecia superficially attached to the substrate, often slightly sunken into it, without stipe, roundish when juvenile and nearly entirely closed at the apex, cupulate when mature, at full maturity with ripped edges; lobes erect when juvenile, effused when mature. Ectal excipulum (epicutis) dark yellow-orange, seldom orange-brown, pseudoparenchymatous, with angular or isodiametric, thick-walled cell agglomerates (textura angularis), cells giving rise to dark brown, thick-walled, multiple septate cylindrical finely warted setae, single or cespitose, dark brown in water and 5% KOH. Asci operculate, cylindrical, eight-spored, monoseriate. Paraphyses slender, single, septate, with gentle orange pigmentation giving rise to the colour of the hymenium, slightly thickened and bent at the tip, arranged in palisade order.
Humaria Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 320 (1870)
Type species: Peziza hemisphaerica F.H. Wigg. 1780
Type designated by Denison (1959)
Diagnosis: Cupulae sparsae, rarius gregariae, sessiles, carnosae, majusculae, extus margineque aut ciliatae setosaeve, aut tomentosae, aut subglabrae, rarius totae glabrae. Disco concavo, demum applanato convexove, discolori. — Asci elongato-cylindracei, 8spori. Sporidia ovata oblongove-ovata, plerumque guttulis 1—3 magnis, continua, hyalina dilutissimeve rubella. Paraphyses filiformes, variae. Plerumque terrestres.
Family: Pyronemataceae
Hydnobolites Tul. & C. Tul.
Ann. Sci. Nat. Bot., sér. 2, 19: 378 (1843)
Type species: Hydnobolites cerebriformis Tul. & C. Tul. 1843
Type designated by Tulasne & Tulasne (1843)
Diagnosis: Peridium vere nullum; substantia carnosa compacta similaris irregularis extus anfractuosa exarata, intus sinubus serpentinis partis, fungi ad superficiem apertis, varie pertusa; sporangia ovato-elliptica inordinate in substantia (extremae filamentorum ex quibus illa constat vesiculae auctae) nidulantia, sporas octo sphaericas reticulato-echinatas foventia. — Fungus globosus basi absorbente, mycelii fibrillis tenuissimis humo adhaerenti praeditus.
Family: Pezizaceae
Hydnocystis Tul. & C. Tul.
G. bot. ital., 1 (7-8): 59 (1845)
Type species: Hydnocystis arenaria Tul. & C. Tul. 1845
Type designated by Tulasne & Tulasne (1845)
Diagnosis: Peridium seu integumentum exterius non discretum nec separabile, crassiusculum, continuum, nusquam interruptum nec aper- tum, superne tantum in centro paulo diminutum, extusque sub-verruculosum. Cavitas interior unica vacua, hujusce paries strato gossypino denso candidissimo sicco, e fibris simplicibus mollibus tenuibus discretis liberis parallele consitis ad centrum fungi vergentibus constanti, tectus; has inter fibras illae asci lineari-oblongi obtusi deorsum abrupte angustati in filum praelongum desinentes, illae paraphyses lineares ascis intermixtis sensim longiores. Thecae sporas octo perfecte sphaericas leves pellucidas glabras in conceptaculi parte superiore dilatata seriatim ordinatas et discretas includentes nec sporas inter constrictae. Fungillus globosus arrhizus, clausus.
Family: Tarzettaceae
Hydnoplicata Gilkey
Mycologia, 46 (6): 783 (1955)
Type species: Hydnoplicata whitei Gilkey 1954
Type designated by Gilkey (1954)
Diagnosis: Ascomatibus lobatis, pucto adjunctionis myceliali destitutis; gleba plicis compactis et irregularibus, nonnumquam anastomosantibus, et fossas longas labyrinthiformesque et cubicula separantibus composita, ascis et paraphysibus in vallum ordinatis vestita; ascis cylindraceis; paraphysibus quam asci longioribus, apicibus in pseudoparenchyma intertextis; sporis uniseriatis, breve ellipsoideis.
Family: Pezizaceae
Hydnotrya Berk. & Broome
Ann. Mag. nat. Hist., ser. 1, 18: 78 (1846)
Type species: Hydnobolites tulasnei Berk. 1844
Type designated by Berkeley & Broome (1846)
Diagnosis: Peridium nullum ; substantia carnosa compacta similis extus anfractuosa exarata, intus sinubus serpentinis magnis fungi ad superficiem apertis varie pertusa filamentis flexuosis mollibus brevibus vestitis. Asci elongati lineares obtusi substantia laxe cellulosa serie unica nidulantes, sporidia octo sphaerica reticulata sed non echinata foventes. Fungi globosi, edules.
Family: Discinaceae
Current definition: Ascomata hypogeous to emergent, irregularly globose to ellipsoid, irregularly lobed to convoluted; hymenophore sequestrate, a ptychothecium, internally hollow or sparsely to highly infolded and chambered; excipular surface shades of brown, glabrous to pubescent. Ascospores brown, globose to ellipsoid or rectangular, nonapiculate to apiculate, thick-walled; surface punctate to nodulose with rounded warts or echinulate at maturity; perispore cyanophilic, contents eguttulate to uniguttulate. Tissues in cross section separated into a medullary excipulum of textura intricata and ectal excipulum of textura angularis or textura prismatica.
Hydnotryopsis Gilkey
University of Calif. Publ. Bot., 6: 336 (1916)
Type species: Hydnotryopsis setchellii Gilkey 1916
Type designated by Gilkey (1916)
Diagnosis: Ascocarp subglobose, compact, more or less lobed and wrinkled, without external opening; cortex sometimes extending deeply into interior; gleba composed of minute, irregular folds, more or less joined, forming complex system of veins separating canals and chambers; hymenium of regular palisade of asci, lining walls of canals and chambers; asci cylindrical, 8-spored; spores ellipsoid, papillose.
Synonym of Sarcosphaera
Hypotarzetta Donadini
Docum. Mycol., 15 (60): 48 (1985)
Type species: Pustularia insignis Berthet & Riousset 1963
Type designated by Donadini (1985)
Diagnosis: Carposoma primum hypogea, stipitata, clausa deinde cupulata ; dein epigaea, cupulata (= concava) vel saepe recurvata-convexa ; pedunculata sed saepe sessilia ; intus hymenium laeve ; extus excipulum furfuraceum. Excipulum medullare supereum a textura intricata compositum ; excipulum medullare inferum a textura fascintricata ; excipulum ectale a textura angulari vel prismatica. Asci non jodo caerulescentes, octospori, cylindrati, operculati saepe aporynchi ; parahyses filiformes, ramosae, cum uninucleatis cellulis ; sporae ellipsoideae uni vel biguttulatae, uninucleatae, laeves vel sublaeves. Omnes cellularum nuclei giemsaphili ; cellulae majorque uninucleatae interdum binucleatae (excipulum, margo, furfuratio).
Synonym of Tarzetta
Imaia Trappe & Kovács
Mycologia, 100 (6): 934 (2008)
Type species: Terfezia gigantea S. Imai 1933
Type designated by Kovács et al. (2008)
Diagnosis: A Terfeziis sporis grandissimis (35–) 42–62 (–69) mm episporam includentibus, epispora amorpha 2–5 mm crassa, canales minutos, sinuosos continenti, et ordinibus DNA divergentibus differt.
Family: Morchellaceae
Current definition: Ascomata globose to ellipsoid or irregular, brown, often cracked at maturity. Peridium of angular cells with thick walls. Gleba composed of brown pockets of asci separated by white veins. Spores globose to subglobose, up to 70 mm long, with a thick, amorphous epispore. DNA sequences divergent from those of Terfezia.
Infundibulum Velen.
Monogr. Discom. Bohem.: 351 (1934)
Type species: Peziza linteicola W. Phillips & Plowr. 1887
Type designated by Eckblad (1968)
Diagnosis: Apoth. mediocria, carnosa, fragilia, primum infundibuli-formia, dein explanata, pileata, margine deflexo, minute crenulato, centro profunde perforata, extus furfuracea, stipite crasso. As. longe cylindrici, truncati, jodo coerulei, par. simpl. fil., apice non dilatatae. Sp. monostichae, ellipticae, eguttulatae, hyalinae.
Synonym of Peziza
Iodomarcelleina Sammut, P. Alvarado & Van Vooren
Cryptog. Mycol., 44 (12): 164 (2023)
Type species: Iodomarcelleina obscura Sammut, P. Alvarado & Van Vooren
Type designated by Sammut et al. (2023)
Family: Pezizaceae
Current definition: Ascomata epigeous, discoid, sessile, dark coloured. Flesh without latex. Asci operculate, with croziers, having a diffusely amyloid wall in iodine solution. Ascospores globose, multiguttulate, smooth. Paraphyses with an external pigment at the top. Ectal excipulum of textura globulosa/angularis. Anamorph unknown. Trophic status unknown.
Iodophanus Korf
in Kimbr., Am. J. Bot., 54 (1): 18 (1967)
Type species: Ascobolus carneus Pers. 1801
Type designated by Korf, in Kimbrough (1967)
Diagnosis: Apothecia minuta, glabra, plerumque pigmentis carotenoideis colorata; asci in iodo caerulescentes; ascospori notis calloso-pecticis ornati, bullulis vel vacuolis evidentibus aliis deficientibus; status conidioferus Oedocephali instar quoad cognitus est.
Family: Pezizaceae
Current definition: Apothecia scattered to gregarious, closed in youth, expanding, becoming plane to convex, roughened with protruding asci, 1 to 2 mm in diam, pallid to yellow or flesh-colored, glabrous; excipulum of two areas, (1) ectal of pallid to pigmented textuia angularis, (2) medullary of textura intricata; asci blued in iodine (slightly stronger at their apices), staining uniformly in congo red, 150-350 µ long, 25-40 µ wide, operculate; ascospores 1 to 2 seriate, elliptical, 8.5-25 × 16-38 µ, finely warted to large (up to 3.0 µ) tack-head markings, without deBary bubbles or other guttules; paraphyses stout, simple or forked, septate, up to 10 µ thick at the apex, with or without oil droplets or pigments. Conidial state where known belonging to the genus Oedocephalum.
Asexual morph of Oedocephalum type.
Iodowynnea Medel, Guzmán & S. Chacón
Mycotaxon, 59: 128 (1996)
Type species: Galactinia auriformis Pat. ex Le Gal 1953
Type designated by Medel et al. (1996)
Diagnosis: A Wynnea affine differt ascis iodinos et sporae verrucosis in vittas in longitudinem porrectas confluentibus.
Family: Pezizaceae
Ionopezia Van Vooren
Ascomycete.org, 12 (4): 187 (2020)
Type species: Peziza gerardii Cooke 1875
Type designated by Van Vooren (2020)
Family: Pezizaceae
Current definition: Ascomata epigeous, discoid or cupulate, sessile, violet-coloured. Flesh without latex. Asci operculate, 8-spored, with or without crozier, wall diffusely amyloid (W type). Paraphyses hyaline, with an external purplish brown to violet pigment located at the top. Ascospores fusoid, multiguttulate, ornamented with longitudinal striations (only seen with a high magnification). Species forming ectomycorrhizas.
Iotidea Clem.
Gen. fung.: 89 (1909)
Type species: Peziza pleurota W. Phillips 1887
Type designated by Clements (1909)
Diagnosis: Asci blue with iodin. Apothecia cleft on one side, ear-like
Synonym of Peziza ?
Jafnea Korf
Nagaoa, 7: 5 (1960)
Type species: Peziza fusicarpa W.R. Gerard 1873
Type designated by Korf (1960)
Diagnosis: Apothecia patellata vel urceolata, 0.5-5(-8)cm. diam., excipulo exteriore e textura prismatica vel angulari, cellulis perpendiculariter ad apothecii faciem exteriorem elongatis. exterioribus brunneis, interioribus hyalinis vel brunneis; pilis superficialibus, brevibus, brunneis, vel in specie una absentibus; hyphae basaleres abundantes, subhyalinae, humi granula amplectentes et pseudostipitem isto modo efficientes; asci operculati, octospori, apicibus in Iodo non caerulescentibus; ascosporae grandae, hyalinae, primum biguttulatae deinde non guttulatae, verrucis minutis vel grandis ornatae. In humo et residuis vegetabilium. Holotypus : Peziza fusicarpa Gerard.
Family: Pyropyxidaceae
Current definition: Apothecia discoid to deep cupulate, 0.5-8 cm. diam.; ectal excipulum composed of cells elongated perpendicularly to the outer surface of the apothecium, outermost cells brown-walled, inner cells hyaline or brown; hairs absent, or if present superficial, short, brown; basal hyphae abundant, nearly hyaline, enmeshing particles of soil to form a pseudostipe or cushion; asci operculate, 8-spored, J-; ascospores large, hyaline, biguttulate in youth, non-guttulate at maturity, marked with small to large warts. On soil and duff.
Kalaharituber Trappe & Kagan-Zur
Mycol. Res., 109 (2): 242 (2005)
Type species: Tuber pfeilii Henn. 1897
Type designated by Ferdman et al. (2005)
Diagnosis: A Terfezia sporis spinis minutissimis dense congestis, ordiniibus DNA abhorrentibus, distributione in Hemisphaerio Meridionali differt.
Family: Pezizaceae
Number of species: 1 fide DoF
Kalapuya M.J. Trappe, Trappe & Bonito
Mycologia, 102 (5): 1059 (2010)
Type species: Kalapuya brunnea M.J. Trappe, Trappe & Bonito 2010
Type designated by Trappe et al. (2010)
Diagnosis: Ascomata hypogaea, stereothecia, subglobosa. Peridium porphyreum vel brunneum, verrucosum, verrucis fissuris tenuibus separatis. Gleba solida, albida, contextu fertili griseo-brunneo maculato. Asci ellipsoidei vel globosi. Sporae ellipsoideae, laeves, juventute hyalinae, maturitate succineae.
Family: Morchellaceae
Kallistoskypha Pfister, Agnello, Lantieri & LoBuglio
Mycol. Progr., 12 (4): 670 (2013)
Type species: Caloscypha incarnata Duvernoy & Maire 1917
Type designated by Pfister et al. (2013)
Family: Caloscyphaceae
Current definition: Apothecial ascomata up to 16 mm diam, hymenium pink, salmon to yellow-cream, often with a short stipe. Outer surface granulose or furfuraceous. Ascospores globose, guttulate. Asci 8-spored, inamyloid, operculate, often with gelatinous granules on the outside. Paraphyses septate, branched only slightly enlarged at the apex, vacuoles and cytoplasm with pigments. Medullary excipulum of several layers of globose to angular cells. Ectal excipulum of globose angular cells arranged in parallel rows, some of the outer cells giving rise to hyaline, often encrusted hairs. Occurring on soil and plant debris around Eucalyptus spp.
Karstenella Harmaja
Karstenia, 9: 20 (1969)
Type species: Karstenella vernalis Harmaja 1969
Type designated by Harmaja (1969)
Diagnosis: Genus Discomycetum, verisimiliter ex ordine Pezizales. Ascocarpus tenuissimus, omnino stricte ad substratum appressus (haud apothecium sensu stricto), probabiliter gymnocarpus. Sporae unicellulatae, satis tenuitunicatae, in KOH hyalinae, in solutione Melzeri subflavidae, in «Cotton Blue« in acido lactico cyanophiloidae. Asci eutunicati, unitunicati, operculati, satis tenuitunicati, tunica ad apicem ejusdem crassitudinis, cylindracei, octospori, in solutione Melzeri haud caerulescentes, sporas per ascostoma leniter obliquum ejectant. Paraphyses recti, simplices, filiformes, in solutione Melzeri haud colorescentes. Excipulum omnino e textura intricata. Subiculum (?) e hyphis tenuibus intricatis, hypharum excipuli dissimilibus.
Family: Karstenellaceae
Current definition: Ascocarp very thin, totally tightly appressed at the substrate (not being a true apothecium), its development probably gymnocarpic. Spores one-celled, rather thin-walled, hyaline in KOH, slightly yellowish in Melzer''s, blue in Cotton Blue (Methyl Blue) in lactic acid. Asci with persistent wall of the unitunicate type, discharging their spores with a puff, operculate, rather thin-walled, apex not differentiated, cylindrical, eight-spored, not blueing with Melzer''s. Excipulum throughout composed of more or less interwoven, filamentous hyphae (textura intricata). Subiculum (?) composed of interwoven, filamentous hyphae of other kind than those of the excipulum.
Kimbropezia Korf & W.Y. Zhuang
Mycotaxon, 40: 272 (1991)
Type species: Kimbropezia campestris Korf & W.Y. Zhuang 1991
Type designated by Korf & Zhuang (1991)
Diagnosis: Ab Peziza structura cyanophilica lentiformi intra parietem operculi asci inclusa differens
Synonym of Peziza
Kompsoscypha Pfister
Mem. N. Y. bot. Gard., 49: 340 (1989)
Type species: Plectania chudei Pat. ex Le Gal 1953
Type designated by Pfister (1989)
Diagnosis: Apothecia parva, minus quam 1 cm diam., plana vel concava; lutea, crocea, vel rubri. Ascis 8-sporis, parietibus crassis. Ascosporis ellipsoideis, eguttulatis vel multiguttulatis. Excipulum medullare e textura intricata; excipulum exterum e textura angulare vel texturam globulosam vergens. Paraphyses filiformes.
In foliis, seminibus, vel lignis tropicarum.
Family: Sarcoscyphaceae
Korfiella D.C. Pant & V.P. Tewari
Trans. Br. mycol. Soc., 54 (3): 493 (1970)
Type species: Korfiella karnika D.C. Pant & V.P. Tewari 1970
Type designated by Pant & Tewari (1970)
Diagnosis: Apothecia coriacea, auriculata, externo excipulo porphyreo brunneo, hymenio-atro. Asci iodo haud tincti; ascosporae hyalinae, ovales vel ellipticae, uni vel biguttulatae; paraphyses simplices vel ramosae, anastomosantes, hyalinae, apice inflato vel angusto stricto.
Family: Sarcosomataceae
Number of species: 1 fide DoF
Kotlabaea Svrček
Česká Mykol., 23: 85 (1969)
Type species: Peziza deformis P. Karst. 1867
Type designated by Svrček (1969)
Diagnosis: Apothecia solitaria usque gregaria, minuta, breviter cylindracea, basi late sessilia, molliter carnosa, aurantiaca, disco plano, anguste marginato, extus nuda, basi hyphis hyalinis instructa sed absque hypothallo typico. Excipulum textura globuloso-angulosa, hyphis marginalibus brevibus, clavatis. Medulla intermixta, plerumque e hyphis utriculosis, irregularibus constat. Hypothecium e cellulis parvis, subglobosis. Asci cylindracei, octospori, membrana non amyloidea. Paraphyses crassae,. apice dilatatae, pigmento aurantiaco impletae. Sporae obtuse ellipsoideae, hyalinae, intus dense granulosae vel minute guttulatae, laeves, membrana subincrassata instructae.
Hab. Ad terram udam.
Synonym of Byssonectria
Labyrinthomyces Boedijn
Bull. Jard. bot. Buitenz., sér. 3, 16 (2): 238 (1939)
Type species: Labyrinthomyces steenisii Boedijn 1939
Family: Tuberaceae
Current definition: Ascocarps hypogeous with a brown tomentose peridium. Gleba with prominent, labyrinthine, mostly hollow chambers that do not open to the surface. Asci cylindric, long-stipitate, mostly 8-spored. Paraphyses much shorter than asci. Spores globose, hyaline, smooth in youth, at maturity ornamented with rounded, irregular warts.
Lamprospora De Not.
Comm. Soc. crittog. Ital., 1 (5): 388 (1863)
Type species: Lamprospora miniata P. Crouan & H. Crouan ex De Not. 1864
Type designated by De Notaris (1863)
Diagnosis: Ascomata exigua, scutellata, sessilia, basi filamentis tenuissimis hyalinis radicantia, extus papillosa, disco planiuscula, tenuiter marginata. Excipulum crassiusculum, cellulis stratosis periphericis coloratis, intimis cum hypothalamicis hyalinis laxioribus. Asci copiosissimi, conferti, e basi tenuata cylindracei leptodermi, 8-spori. Paraphyses filiformes, apice colorato, nonnihil incrassatæ. Sporidia hyalina, sphæroidea, 8/500. mm. Diametro æquantia, episporio elegantissime reticulato, areolis hexagonis prædita. Sulla terra, tra i muschi, nelle pasture di Val Intrasca, autunno 1862, rara.
Family: Pyronemataceae
Lasiobolidium Malloch & Cain
Can. J. Bot., 49 (6): 853 (1971)
Type species: Lasiobolidium spirale Malloch & Cain 1971
Type designated by Malloch & Cain (1971)
Diagnosis: Ascogonia convoluta. Ascocarpae subglobosae vel globosae, luteo-brunneae, appendiculatae, nonostiolatae. Asci irregulariter dispositi, cum agoli, octospori, evanescentes, nonstipitati vel brevi stipitati, nonamyloidei. Ascosporae ellipsoideae, unicellares, hyalinae, laeves, sine pora. Conidia ignota.
Family: Pyronemataceae
Current definition: Ascomata cleistothecial or apothecial, small, sessile, subglobose, slightly cupuliform or discoid, whitish, pale greyish or yellow-brown, with an external surface covered by brown hairs. Margin hairy on the apothecial species. Ectal excipulum of textura globulosa/angularis. Hairs ± superficial or deeply rooted, septate, with a simple base, sometimes multifurcate, long, upward-pointing, straight or helically coiled. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 4- or 8-spored. Ascospores ellipsoid, hyaline, eguttulate, but sometimes containing small granules usually disappearing in mature ascospores, smooth or verrucose with isolated warts. Species saprobic, coprophilous or not. Asexual morph unknown.
Lasiobolus Sacc.
Bot. Zbl., 18: 220 (1884)
Type species: Peziza equina O.F. Müll. 1778
Type designated by Clements & Shear (1931)
Diagnosis: Lasiobolus Sacc. (Ascophanus Boud. p. p.) Ascomata scutellato-cupulata, distincte setosa. Asci et sporidia Ascophani.
Family: Ascodesmidaceae
Lasiocupulina Van Vooren & M. Vega
Ascomycete.org, 10 (6): 221 (2018)
Type species: Lasiocupulina mediterranea Van Vooren & M. Vega 2018
Type designated by Van Vooren & Vega (2018)
Diagnosis: Differs from Cupulina by the presence of a hairy margin and its molecular data.
Family: Pyronemataceae
Lathraeodiscus Dissing & Sivertsen
Mycologia, 80 (6): 832 (1989)
Type species: Lathraeodiscus arcticus Dissing & Sivertsen 1989
Type designated by Dissing & Sivertsen (1989)
Diagnosis: Carposomata parva, sessilia, disciformia, marginata, extra glabra vel furfuracea, omnino fuscida purpureo vel vinaceo affecta. Excipulum exterius tenue, e cellulis hyphoidibus formatum ad marginem versus claviformibus, contento cinereo-violaceo, homogeneo vel granulari. Excipulum medullare e cellulis globularibus vel angulatis formatum. Subhymenium manifestum. Asci operculati, cylindrici, non amyloides, contento pallide cinereo-violaceo, ad bases elongati, pleurorhynchi. Paraphyses rectae, septatae, ramificatae, contento cinereo-violaceo, homogeneo vel granulari. Ascosporae uniseriatae, ellipsoides, guttulas singulas continentes, colore nullo, ornamento cyanophilo.
Family: Pyronemataceae
Lazuardia Rifai
Mycotaxon, 31 (1): 241 (1988)
Type species: Peziza lobata Berk. & M.A. Curtis 1868
Type designated by Rifai (1988)
Diagnosis: Apothecia gregaria, mediocria, sessilia, denique discoidea, disco caerulescentes, extra tomentosa ex pilis hyalinis septatis composita. Excipulum ectale e cellulis angularibus (textura angularis), excipulum medullare ex hyphis septatis (textura intricata) compositum. Asci cylindracei, apice operculati jodo haud tincti, octospori. Ascospori uniseriati, hyalini, globosi, ornati, uniguttulati. Paraphyses septatae, rectae, filiformes.
Family: Pulvinulaceae
Legaliana Van Vooren
Ascomycete.org, 12 (4): 188 (2020)
Type species: Peziza badia Pers. 1800
Type designated by Van Vooren (2020)
Family: Pezizaceae
Current definition: Ascomata epigeous, cupulate or discoid, sessile, dark coloured (brown, reddish brown, olivaceous brown, purplish brown, blackish purple). Flesh without coloured latex, but sometimes with a watery juice. Asci operculate, 8-spored, with crozier, wall diffusely bluing in an iodine solution, more or less intensely at the top (WT type). Paraphyses containing a brownish or olive-brown vacuolar pigment, and often showing an external brown pigment at the top. Ascospores biguttulate, sometimes with an oil drop larger than the other, more rarely uniguttulate, ornamented with irregular warts, but often with elongated warts or crests, sometimes forming a network or an incomplete reticulum. Ectal excipulum of textura globulosa/angularis, with large cells. Species saprobic or ectomycorrhizal.
Lepidotia Boud.
Bull. Soc. mycol. Fr., 1: 101 (1885)
Type species: Peziza hispida Quél. 1879
Type designated by Eckblad (1968)
Diagnosis: Réceptacles sensiblement pédiculés ou obconiques, à squames triangulaires submembraneuses. Spores elliptiques, sans sporidioles.
Family: Pezizaceae
Leptopodia Boud.
Bull. Soc. mycol. Fr., 1: 99 (1885)
Type species: Helvella elastica Bull. 1785
Type designated by Boudier (1885)
Diagnosis: Réceptacles infléchis de plusieurs côtés sur le stipe, entiers ou profondément lobés, à bords libres ou soudés par places au pédicule. Spores elliptiques avec une grosse sporidiole centrale accompagnée de quelques granulations plus petites à chaque extrémité. Réceptacles entiers ou à peine lobés, pédicule relativement grêle.
Synonym of Helvella
Leucangium Quél.
Compt. Rend. Assoc. Franç. Avancem. Sci., 11: 404 (1883)
Type species: Leucangium ophthalmosporum Quél. 1883
Diagnosis: Peridium globuleux (0m,03–4), mince, inséparable, finement grenelé, souvent crevassé, couvert de fins poils soyeux et bruns, surtout en dessous, châtain, puis d''un beau noir mat. Glèbe charnue, ferme, grenue (celluleuse à la loupe), blanc-crème, violette extérieurement, réticulée de blanc et pointillée. Thèque globuleuse à 6 spores (rarement 4 ou 8). Spore oculiforme (0mm,06-8), ocellée, faiblement chagrinée, olivâtre. Odeur de melon. (pl. XII, fig. 2.)
Family: Morchellaceae
Leucopezis Clem.
Gen. Fungi: 175 (1909)
Type species: Leucopezis excipulata Clem. 1909
Type designated by Clements (1909)
Diagnosis: Neottiopezis eciliata
Synonym of Humaria ?
Leucoscypha Boud.
Bull. Soc. mycol. Fr., 1: 104 (1885)
Type species: Peziza leucotricha Alb. & Schwein. 1805
Type designated by Le Gal (1957)
Diagnosis: Poils blancs, spores verruqueuses ou réticulées.
Family: Pyronemataceae
Loculotuber Trappe, Parladé & I.F. Alvarez
Mycologia, 84 (6): 927 (1993)
Type species: Terfezia gennadii Chatin 1896
Type designated by Trappe et al. (1993)
Diagnosis: A Tuber loculis casis necnon ascis stipitato-ellipsoides vel clavatis in hymeniis differt.
Synonym of Tuber
Lotinia Pérez-Butrón, Fern.-Vic. & P. Alvarado
in Crous et al., Persoonia, 35: 325 (2015)
Type species: Lotinia verna Pérez-Butrón, Fernández-Vicente & P. Alvarado 2015
Type designated by Pérez-Butrón et al. (2015)
Family: Pyronemataceae
Current definition: Ascomata discoidal, brownish, with the external surface covered with flexuose hairs from the base to the top. Ectal excipulum arranged as a textura prismatica. Asci 8-spored, operculate, not amyloid. Ascospores smooth, not guttulated, globose to ellipsoid.
Luciotrichus R. Galán & Raitv.
Czech Mycol., 47 (4): 272 (1995)
Type species: Luciotrichus lasioboloides R. Galán & Raitv. 1995
Type designated by Galán & Raitviir (1995)
Diagnosis: Apothecia sessilia, turbinata vel cupulata, pallide colorata, extus longe rigidipilosa. Pili hyalini, conici, valde crasso-tunicati, multiseptati. Asci operculati, cylindraceo-clavati, inamyloidei, octospori. Sporae ellipsoideae vel late ellipsoideae, non guttulatae, aliquando "de Bary bubble" praeditae, verrucosae. Paraphyses filiformes, apicibus rectis vel subcurvatis.
Mycoarctium Jain & Cain similis, ascis operculatis differt.
Family: Ascodesmidaceae
Luteoamylascus Cabero, P. Alvarado & G. Moreno
Mycol. Progr., 15 (4): 3 (2016)
Type species: Luteoamylascus aculeatus Cabero, P. Alvarado & G. Moreno 2016
Type designated by Cabero et al. (2016)
Diagnosis: A combination of smooth peridium, amyloid asci, and globose spores ornamented with large spines. Genetic profile differs from all other sequenced, described Pezizaceae
Family: Pezizaceae
Macropodia Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 331 (1870)
Type species: Macropodia macropus (Pers.) Fuckel 1870
Type designated by Fuckel (1870)
Diagnosis: Cupulae magnae, sparsae, carnosae, plano-hemisphaericae, abrupte longe stipitatae, totae hirto-pulverulentae, disco vix discolori. Asci longissimi, cylindracei, 8spori. Sporidia in asci superiori parte oblique monosticha, ovato-ellipsoidea, uniguttulata, hyalina, laevia, continua. Paraphyses filiformes adsunt. Terrestres autumnalesque.
Synonym of Helvella
Macroscyphus Nees ex Gray
Nat. Arr. Brit. Plant., 1: 671 (1821)
Type species: Macroscyphus macropus (Pers.) Gray 1821
Diagnosis: Long-funnel. Thallus cuplike, soft, cellular, rather fibrous ; surface scaly or slightly bristly ; sporidia with 6 or 8 sporae in each, in a single row.
Synonym of Helvella
Malvipezia Van Vooren
Ascomycete.org, 12 (4): 188 (2020)
Type species: Peziza howsei Roze & Boud. 1880
Type designated by Van Vooren (2020)
Family: Pezizaceae
Current definition: Ascomata epigeous, cupulate or discoid, sessile, with pale mauve, purple or violet colours. Flesh without latex. Asci operculate, 8-spored, with crozier, wall diffusely amyloid in an iodine solution except at the top where the reaction is intense and forms a ring (WTR type). Paraphyses containing a brown vacuolar pigment. Ascospores biguttulate, ornamented with isolated, rounded warts, more rarely with irregular and coalescent warts. Ectal excipulum of textura globulosa with medium-sized to large cells. Species saprobic.
Marcelleina Brumm., Korf & Rifai
Persoonia Suppl., 1: 233 (1967)
Type species: Ascobolus persoonii P. Crouan & H. Crouan 1867
Type designated by Brummelen (1967)
Diagnosis: Apothecia sessilia, interdum ad basin paullo constricta, terricola. Receptaculum carnosum, molle, sat parvum, demum expansum. Discus e piano convexus, violaceus, brunneo-violaceus, brunnescens aut nigrescens, pigmento carotenoideo miniato aut rubro destitutus. Asci cylindracei, operculati, pariete iodo non caerulescente. Ascosporae globulosae, ornatae aut laeves. Paraphyses tenues, apice incrassatae saepe curvatae vel curvulae.
Family: Pezizaceae
Mattirolomyces E. Fisch.
Nat. Pflanzenfam. (Ed. 2), 5bVIII: 39 (1938)
Type species: Choiromyces terfezioides Mattir. 1888
Family: Pezizaceae
Maublancomyces Herter
Rev. Sudam. de Botanica, 8: 161 (1950)
Type species: Helvella gigas Krombh. 1834
Type designated by Herter (1950)
Diagnosis: Sporae fusiformes, apicibus verruca ornatae, asperae vel subtilissime irregulariter reticulatae, 30-45 micra long., guttula magna centrali et saepius duabus minoribus polaribus praeditae.
Family: Discinaceae
Current definition: Ascomata epigeous, substipitate to stipitate; hymenophore cupulate to discoid or convex and lobed, saddle-shaped or cerebriform; hymenium yellow-ochre, buff, yellow–brown, brown, or reddish-brown; stipe broadly cylindric, irregular, or fluted, chambered, white to yellow–brown. Ascospores hyaline, ellipsoid to fusoid, apiculus broadly rounded or blunt, sometimes knobbed; surface rugulose to somewhat reticulate at maturity; perispore cyanophilic; contents uni- or triguttulate.
Melascypha Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Peziza melaena Fr. 1822
Type designated by Boudier (1885)
Diagnosis: Thèques très longues et très grêles, à base flexueuse pénétrant dans le parenchyme; paraphyses souvent très rameuses (Longithèques). Espèces à hyménium d’un beau noir; paraphyses à ramifications moins pointues. Espèces glabres, pédiculées. Spores rondes.
Synonym of Pseudoplectania
Melastiza Boud.
Bull. Soc. mycol. Fr., 1: 106 (1885)
Type species: Humaria miniata Fuckel
Type designated by Seaver (1928)
Diagnosis: Poils courts et obtus; spores avec ou sans sporidioles. Spores à deux sporidioles ou gouttelettes. Espèces terrestres ou carbonicoles; couleur rouge ou orangée. Espèces à spores réticulées ; terrestres.
Family: Pyronemataceae
Microeurotium Ghatak
Ann. Bot. Lond., 50: 860 (1936)
Type species: Microeurotium albidum Ghatak 1936
Type designated by Ghatak (1936)
Family: Incertae sedis
Number of species: 1 fide DoF
Micronematobotrys Xiang Sun & L.D. Guo
Mycol. Progr., 9 (4): 569 (2010)
Type species: Micronematobotrys verrucosus Xiang Sun & L.D. Guo 2010
Family: Pyropyxidaceae
Number of species: 1
Current definition: Conidiophores micronematous, pale brown, smooth or with ornamentation, unbranched. Conidiogenous cells holoblastic, polyblastic, integrated, intercalary, hyaline to pale brown, inflated laterally, thin-walled, minute denticulate. Conidia solitary, simple, pale brown, verrucose to capitate, 1-celled, ellipsoidal, obovoid, clavate, straight or slight curved, base truncate, apex obtuse, eguttulate.
Microstoma Bernstein
Nova Acta Acad. Caes. Leop.-Carol. Nat. Cur. Dresden, 23: 649 (1852)
Type species: Microstoma hiemale Bernstein 1852
Type designated by Bernstein (1852)
Diagnosis: Cupula subglobosa, ostiolo exiguo poriformi, demum lacero-dehiscens l. dilabens, rhizomali insidens. Asci immersi, paraphysibus tenuibus subramosis succo colorato repletis interstincti.
Family: Sarcoscyphaceae
Midotis Fr.
Elench. fung., 2: 29 (1818)
Type species: Midotis lingua Fr. 1828
Type designated by Fries (1828)
Diagnosis: CHAR. Hymenium inferum ! laevissimum (exsiccatione rimosum) a pileo discretum et separabile. Asci immersi, longi. — Receptaculum carlilagineum, forma omnino peculiari distinctum.
Obs. Forma longe aurita Midae aures in mentem revocavit indeque nomen desumsi. Hymenium Pezizarum quarundam, v. c. P. vesiculosae, simile, sed ob hymenium inferum ad Auricularinos referenda.
Family: Helvellaceae
Miladina Svrček
Česká Mykol., 26 (4): 213 (1972)
Type species: Peziza lecithina Cooke 1876
Type designated by Svrček (1972)
Diagnosis: Apothecia haud magna late sessilia, subdisciformia, crasse carnosa, marginata, extus margineque subglabra, basi solum hyphis affixa. Excipulum textura globulosa, e cellulis globosis, maioribus, tenuiter tunicatis, pigmente aurantiaco impletis, margine parteque exteriore cum hyphis cylindraceoclavatis. Asci tenuiter tunicati, operculati, cylindracei, octospori, membrana non amyloidea. Paraphyses rectae, copiosae, simplices, apice subclavatae, pigmento aurantiaco impletae, vi iodi virescentes. Ascosporae ellipsoideae, tenuiter tunicatae, hyalinae, guttulatae, verrucosae, verrucis cynophilis. Lignicola, hygrophila.
Family: Pyronemataceae
Asexual morph of Actinosporella type.
Mitrophora Lév.
Ann. Sci. Nat., Bot., sér. 3, 5: 249 (1846)
Type species: Mitrophora semilibera (DC.) Lév. 1846
Diagnosis: Receptaculum carnosum conicum vel campanulatum alveolatum, stipitatum, e medio ad marginem a stipite discretum. Thecæ cylindricæ, octosporæ paraphysibus filiformibus immixtæ. Sporæ ovatæ continuæ.
Synonym of Morchella
Monascella Guarro & Arx
Mycologia, 78 (5): 869 (1986)
Type species: Monascella botryosa Guarro & Arx 1986
Type designated by Guarro & Arx (1986)
Diagnosis: Mycelium expandum, ex hyphis septatis, dilute ochraceis vel hyalinis; ascomata ex ascogoniis clavatis et antheridiis spiralibus oriunda, superficialis, discreta vel confluentia, nuda, 80–160 µm; paries basilis ex hyphis hyalines vel brunneis composita; asci botryosi, clavati vel saccati, stipitati, unitunicati, inoperculati, octosporibus, 30–42 × 14–17 µm; ascosporae ellipsoideae, glabrae, hyalinae, biguttulatae, 9–11 × 6–7 µm; paraphyses nullae; anamorphosis ignota, conidia absunt.
Family: Otideaceae
Number of species: 1 fide DoF
Moravecia Benkert, Caillet & Moyne
Z. Mykol., 53 (1): 140 (1987)
Type species: Humaria calospora Quél. 1885
Type designated by Benkert (1987)
Diagnosis: Apothecia minuta, sessilia, paulum immersa, primo globosa clausa deinde cupuliformia, margine sterile pallida et denticulata. Hymenio rubro-aurantia. Excipulum textura angularis compositum, margine textura porrecta compositum. Asci cylindracei, octospori, apice jodo non caerulescentes. Ascospori ellipsoidei, hyalini, juvente multi guttulati, deinde non guttulati, sed ex de Barry guttulatis, reticulati, sine apiculatis. Paraphyses septatae, rectae vel non multum curvatae, apice subclavatae et aurantiaceae coloratae.
Family: Pyronemataceae
Morchella Dill. ex Pers.
Neues Mag. Bot., 1: 116 (1794)
Type species: Phallus esculentus L. 1753
Type designated by Persoon (1794)
Diagnosis: Pileo conico reticulato-lacunoso.
Family: Morchellaceae
Muciturbo P.H.B. Talbot
Mycol. Res., 92 (1): 95 (1989)
Type species: Muciturbo reticulatus P.H.B. Talbot 1989
Type designated by Warcup & Talbot 1989
Diagnosis: Apothecia parva, hypogaea, superne hemisphaerica vel lenticulato-turbinata, infra compressa vel concava, saepe per acumen centrale obconicum affixa vel demum vel demum excavata, laevia vel lobata, initio firma et albida demum violaceo-lilacina vel atrobrunnea, hymenio supra superficiem externam. Excipulum medullosum e textura globulosa vel textura angulari. Asci e pedo oriundo immaturi breviter amyloidei ascosporis in turma irregulari prope apicem, maturi subcylindrici parietibus tenuis et omnino non amyloidei, evidenter sine structura apicale dehiscentia consociata, deliquescentes ubi maturi ut sporas liberarent, plerumque ascosporis 8, rare minus. Ascosporae uniseriatae uninucleatae, sphaericae, hyalinae, tum violaceo-lilacinae, demum atrobrunneae vel atrae, semper ornatae, haud amyloideae. Paraphyses hyalinae vel pallido-stramineae, sparsim ramosae ad basim et interdum omnino per longitudinem totam, ad apices leviter parvae, tenuitunicatae vel tunicis gelatinosis crassis, septis frequentibus, extensae super ascis ut epithecium formarent, gelatinescentes et demum deliquescentes in muco qui ascosporas tolleret.
Synonym of Ruhlandiella
Mycoarctium K.P. Jain & Cain
Can. J. Bot., 51 (2): 305 (1973)
Type species: Mycoarctium ciliatum K.P. Jain & Cain 1973
Family: Incertae sedis
Number of species: 2 fide DoF
Mycoclelandia Trappe & G.W. Beaton
Trans. Br. mycol. Soc., 83 (3): 536 (1984)
Type species: Clelandia arenacea Trappe 1979
Type designated by Trappe (1979) under the generic illegitimate name Clelandia.
Diagnosis: Ascocarpia exsiccata ± 2 cm in diam, profunde sulcata. Peridium glabrum, pallide fulvum, arena adherenti velatum. Gleba loculis vacuis labyrinthinis, hymeniis limitatis. Asci cylindrici vel saccati, octospori, valde amyloidei. Sporae late ellipsoideae vel subglobosae, 10-12 × 8-10 µm in diam, dispersae, inferiores quam asci. Holotypus: ADW 15885.
Family: Pezizaceae
Number of species: 2 fide DoF
Mycogalopsis Gjurašin
Acta bot. Inst. bot. Zagreb, 1: 9 (1925)
Type species: Mycogalopsis retinospora Gjurašin 1925
Family: Pyronemataceae
Myrmecocystis Harkn.
Proc. Calif. Acad. Sci., Ser. 3, Bot., 1: 269 (1899)
Type species: Myrmecocystis cerebriformis Harkn. 1899
Type designated by Clements & Shear (1931)
Diagnosis: Fungus minute, irregular, lobed or gibbous, verrucose; gleba chambered by an irregularly stellate cavity not communicating with the exterior; asci subglobose or somewhat elongate, 8-spored; spores rough.
Family: Pyronemataceae
Nanoscypha Denison
Mycologia, 64 (3): 617 (1972)
Type species: Cookeina tetraspora Seaver 1925
Type designated by Denison (1972)
Diagnosis: Apothecia cupularia vel patellaria, stipitata vel sessilia, modica vel minuta, 1-10 mm lata ; excipulum carnosum vel coriaceum; duobus stratis compositum, strato interiore filamentosa, strato exteriore membranaceum e cellulis subrectangularibus constans ; hymenium pallidum vel coloratum, luteum vel rubrum ; extra simile colore, glabrescente ; asci suboperculati, cylindracei, in basim sensim angustati, non simul maturescentes ; opercula eccentrici ; sporae ellipsoideae vel leviter curvatae, subcymbiformae, laeves, 1-2 guttis adfectae ; paraphyses tenues cylindraceaque, septatae. Plantae ad ramulos et folia marcida, clima calido vigentes.
Family: Sarcoscyphaceae
Current definition: Apothecia small, less than 1 cm broad, cup-shaped to discoid or subturbinate, substipitate to stipitate or sessile, centrally attached, less frequently eccentric; texture firm-fleshy to leathery when fresh, brittle and corky when dry; stipe, where present, similar in color and texture to the cup, slender; hymenium concave to discoid, usually some shade of orange but varying from yellow to red, often translucent; exterior whitish or yellowish, glabrous to glabrescent, becoming strongly wrinkled when dry; ectal excipulum sharply differentiated from the medullary excipulum, textura angularis to textura epidermoidea consisting of several layers of large, 5-30 µ, pseudoparenchymatous cells with their long axes perpendicular to the exterior, sometimes with the outermost layer of cells smaller, 1-5 µ, and with thickened walls; medullary excipulum well developed, 40-200 µ or more thick, textura intricata; asci suboperculate with a lateral operculum, iodine negative, less than 300 µ long, maturing seriatim, cylindrical to clavate with a long tapered, and often crooked, base, 4-spored or with a mixture of 8-, 6-, and 4-spored asci; ascospores mostly unequal-sided ellipsoidal (ellipsoidal and symmetrical in N. macrospora), smooth, containing two conspicuous oil drops; paraphyses filiform to subclavate, not anastomosing to form a network about the asci. On leaves, twigs, and small fragments of wood in the tropics.
Asexual morph of Mollardiomyces type.
Napomyces Setch. ex Clem. & Shear, nom. illeg.
Gen. Fungi: 333 (1931)
Type species: Napomyces gardneri (Setch.) Clem. & Shear 1931
Synonym of Daleomyces
Neogyromitra S. Imai
Bot. Mag., Tokyo, 46: 174 (1932)
Type species: Morchella caroliniana Bosc 1811
Type designated by Imai (1932)
Diagnosis: Genus affine Helvellae sed sporae magnae, subfusiformae, apiculatae et sculpturae.
Family: Discinaceae
Current definition: Ascomata epigeous, substipitate to stipitate; hymenophore cupulate to discoid or convex and irregularly lobed to cerebriform; hymenium rugose, orange-brown to red-brown or purple-brown; stipe broadly cylindric, fluted, or lacunose, white. Ascospores hyaline, ellipsoid to subfusoid, with multiple, prominent, blunt apiculi; surface with a coarse, regular, widely-spaced reticulum at maturity; perispore cyanophilic; contents uniguttulate or triguttulate.
Neottiella (Cooke) Sacc.
Syll. fung., 8: 190 (1889)
Type species: Peziza albocincta Berk. & M.A. Curtis 1875
Type designated by Eckblad (1968)
Diagnosis: Cooke Mycogr. p. 261, ex max. parte [1879] (Etym. neottia nidus). — Ascoma sessile, patellare v. cupulatum, carnosum, sat minutum, pilis tenuibus hyalinis albo-tomentosum ; discolaete colorato vel albido. Asci cylindracei, octospori, paraphysati. Sporidia ellipsoidea v. oblonga, hyalina. — Pyromemati affinis.
Family: Pyronemataceae
Neournula Paden & Tylutki
Mycologia, 60 (6): 1160 (1969)
Type species: Neournula nordmanensis Paden & Tylutki 1969
Type designated by Paden & Tylutki (1969)
Diagnosis: Apothecia stipitata, urceolata vel cupularia; margo crenata vel stellata; hymenium pallidum vel purpureum; externa pallida subbrunnea vel brunnea-purpurea; excipulum medullare e textura intricata; excipulum exterius e textura angulari; pili breves; asci suboperculati, cylindracei, in basim obtusi, in iodo non caerulei, non simul maturescentes; opercula eccentrica: ascosporae ellipsoideae vel oblongateae, verrucoseae, hyalineae; paraphyses tenues, ramosae, anastomosantes
Family: Chorioactidaceae
Current definition: Apothecia stipitate, thin-fleshed, leathery, urceolate to goblet-shaped; margin crenate to stellate; hymenium light colored to dull purplish; exterior pale brown to purplish brown; asci suboperculate, cylindrical, with blunt, occasionally lobed bases, connected to the hypothecium by a narrow hypha, not maturing simultaneously, not bluing in iodine; opercula eccentric; ascospores elliptical or oblong, hyaline, at first smooth, becoming verrucose, the ornamentation staining in heated cotton blue; paraphyses narrow, branched, septate, anastomosing ; medullary excipulum of textura intricata, without an evident gelatinous layer; ectal excipulum of textura angularis, the outermost cells giving rise to short, scattered hairs consisting of individual hyphae.
Nothoamylascus Healy & M.E. Sm.
Persoonia, 51: 145 (2023)
Type species: Nothoamylascus erubescens Healy & M.E. Sm. 2023
Type designated by Healy et al. (2023)
Family: Pezizaceae
Number of species: 1
Current definition: Ascoma a ptychothecium, hypogeous. Excipulum composed of textura angularis or textura globulosa, with large isodiametric to ovoid cells, intermixed with oleiferous hyphae. Gleba composed of asci and paraphyses in a palisade. Asci cylindrical, evenly amyloid in Melzer’s reagent. Ascospores globose, ornamented with truncated spines. Mitotic spore mats in small clusters, epigeous on soil or hypogeous, white with pinkish brown areas, spore mass pink when young but yellow at maturity, mitotic spores nearly smooth to minutely warted, globose to subglobose.
Nothojafnea Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 93 (1968)
Type species: Nothojafnea cryptotricha Rifai 1968
Type designated by Rifai (1968)
Diagnosis: Apothecia gregaria, minuta vel mediocria, brunnea, subsessilia, cupulata, extra hirsuta. Pili rigidi, hyalini, crasse tunicati, septati. Excipulum ectale e cellulis angularibus (textura angulari), excipulum medullare ex hyphis septatis compositum. Asci cylindracei, octospori, apice jodo non caerulescentes. Ascospori ellipsoidei, hyalini, minute verrucosi, guttulati. Paraphyses septatae, apice incrassatae, brunneae. Hab. ad terram.
Family: Pyronemataceae
Ochotrichobolus Kimbr. & Korf
Mycotaxon, 17: 326 (1983)
Type species: Ochotrichobolus polysporus Kimbr. & Korf 1983
Type designated by Kimbrough & Korf (1983)
Diagnosis: Differing from Lasiobolus by septate hairs, from Trichobolus by operculate asci, and from both genera by having hairs present only at the margin and by forming buttresses at the bases of the hairs.
Family: Ascodesmidaceae
Ochromitra Velen.
Monogr. Discom. Bohem.: 391 (1934)
Type species: Helvella sphaerospora Peck 1875
Type designated by Eckblad (1968)
Diagnosis: Pileo ascomatis maximo, campanulato, lobato, cerebriformi, cavo, libero. Stipite crasso, cavo, lacunoso-venoso. Ascis cylindraceis, obtusis, par. simpl. filif., apice incrassatis. Sporis perfecte globosis, hyalinis, laevibus, eguttulatis, minoribus.
Genus proximam Gyromitrae, sed sporis globosis, eguttulatis, stipite lacunoso diversum. Hactenus duas species continens, unam americanam (O. sphaerosporam Peck), alteram europaeam (O. Gabretae Kavina). O. sphaerospora ab autors in paludosis silvaticis Indianlake Amer. bor. Indicator. Seaver: Vermont to Wisconsin and Manitoba.
Synonym of Pseudorhizina
Octospora Hedw.
Descr. micr.-anal. musc. frond., 2: 4 (1789)
Type species: Octospora leucoloma Hedw. 1789
Type designated by Korf (1954)
Diagnosis: Truncus simplicissimus, superficie terminatus fertili concava, plan, reflexa: de qua eriguntur, Thecae seminales membranulosae, elongatae, numerosissimae, utplurimum stupa filamentosa cinctae; utrisque inter se non cohaerentibus, seminibus octo foetae
Family: Pyronemataceae
Octosporella Döbbeler
Nova Hedwigia, 31 (4): 827 (1980)
Type species: Nectria jungermanniarum P. Crouan & H. Crouan 1867
Type designated by Döbbeler (1980)
Diagnosis: Ascocarpia circa 300-500 µm longa, ovoidea ad paene doliiformia, dilute flavida vel aurantiaca, glabra aut setifera, orificio parvo. Paries ascocarpiorum cellulis irregularibus magnisque. Paraphyses filiformes. Asci operculati, plerumque cylindracei vel claviformes, (4-)8spori; J–. Sporae usque ad 40 µm longae et 14 µm latae; unicellulares, ellipsoideae, incoloratae, episporio laevi vel aspero, guttis munitae. Hyphae crassae et crassitunicatae, non coloratae, superficialiter crescentes, appressoriis characteristicis haustoriisque praeditae.
Habitat parasitice in Hepaticis foliaceis.
Family: Pyronemataceae
Octosporopsis U. Lindem. & M. Vega
Z. Mykol., 80 (2): 566 (2014)
Type species: Humaria nicolai Maire 1926
Type designated by Lindemann & Vega (2014)
Family: Pyronemataceae
Current definition: Apothecia 2-7 mm in diameter, sessile, at first more or less obconical, later becoming disc-shaped. Hymenium pale yellow, salmon pink to orange. Margin at first slightly bulged, later more or less membranous, with hyaline hairs; the flanks of the apothecium also covered with hairs. Asci 8-spored, operculate, not amyloid, base pleurorhynchous. Ascospores hyaline, broadly ellipsoid to broadly subfusoid, at maturity thin-walled, with minute refractive light brownish droplets, mostly binucleate, later becoming thick-walled and droplets accumulating in the centre. Paraphyses filiform, apically more or less swollen, terminal cell with faintly to distinctly yellowish homogeneous content. Excipulum two-layered: medullary excipulum of textura intricata/subepidermoidea, composed of thin-walled hyaline cells; ectal excipulum as textura globulosa/angularis, composed of hyaline, thick-walled, isodiametric cells. Hairs hyaline, thick-walled, walls one-layered, multi-septate.
Oedocephalum Preuss
Linnaea, 8: 131 (1851)
Type species: Oedocephalum elegans Preuss 1851
Diagnosis: Hyphasma repens, septatum; stipes erectus, septatus, simplex, supra capitulo homogeneo vesicaeformi ornatus et sporidifero. Sporae simplices, capitulo subradiatim insertae; nucleo farcto.
Family: Incertae sedis
Number of species: 9 fide DoF
Orbicula Cooke
Handb. Brit. Fungi, 2: 926 (1871)
Type species: Orbicula cyclospora Cooke 1871
Type designated by Cooke (1871)
Diagnosis: Perithecia between membranaceous and carbonaceous, reticulated, seated on a distinct mycelium; ostiolum obsolete; asci cylindrical; sporidia subglobose, hyaline or coloured; paraphyses simple or branched (Fig. 401.)
Family: Pseudombrophilaceae
Orcadia G.K. Sutherl.
Trans. Br. mycol. Soc., 5 (1): 151 (1915)
Type species: Orcadia ascophylli G.K. Sutherl. 1915
Family: Incertae sedis
Number of species: 1 fide DoF
Ostracoderma Fr.
Syst. orb. veg., 1: 150 (1825)
Type species: Ostracoderma institale Fr. 1825
Type designated by Fries (1825)
Diagnosis: Peridium rotundato-complanatum, crustaceum, fragile, glabrum, in medio denudatum. Sporidia coacervata. Thallus nullus. Terrestria. Flocci nulli.
Family: Pezizaceae
Number of species: 3 fide DoF
Otidea (Pers.) Bonord.
Handb. Allgem. mykol.: 205 (1851)
Type species: Peziza onotica Pers. 1801
Family: Otideaceae
Oviascoma Y.J. Yao & Spooner
Mycol. Res., 100 (1): 102 (1996)
Type species: Lamprospora crechqueraultii var. paludosa Dennis 1956
Diagnosis: Apothecia solitaria vel gregaria, albida, in sicco brunneola, primum cylindrica demum ovoidea vel obovoidea vel globosa. Discus valde convexus, laevis. Receptaculum emarginatum, glabrum, ad basin angustatum. Excipulum e textura globulosa compositum; muri cellularum tenues; cellulae marginem versus elongatiusculae vel clavatae. Asci operculati, cylindrici, I-. Ascosporae unicellulares, vel subglobosae, spinis ornatae. Paraphyses filiformes, septatae.
Family: Pyronemataceae
Current definition: Apothecia solitary or gregarious, whitish when fresh, brownish-orange to brown after draying; cylindric at fist, becoming ovoid or obovoid to almost globose. Disc strongly convex, smooth. Receptacle emarginate, deeply cupulate, outer surface glabrous, attached to the substratum at a narrow base. Excipulum a textura globulosa, cells thin-walied, broadly ellipsoid to subglobose, marginal cells slightly elongate or clavate. Asci operculate, cylindric, I-. Ascospores unicellular, colourless, globose to subglobose, ornamented with spines. Paraphyses filiform, flexuous.
Pachyella Boud.
Hist. Class. Discom. Eur.: 50 (1907)
Type species: Peziza barlaeana Bres. 1898
Type designated by Le Gal (1953)
Diagnosis: Le réceptacle est toujours épais, peu cupuliforme et plutôt étalé chez les grandes espèces, en coussinet chez les petites. Les thèques sont octospores et bleuissant très légèrement chez plusieurs espèces et pas sensiblement chez d''autres. Les spores sont largement ellipsoïdes, avec deux sporidioles accompagnées de granulations nombreuses. Ce sont des Champignons terrestres ou poussant sur les bois pourris. Les spores ressemblent à celles des Sarcosphaera ; l''aspect général est celui des Plicaria dont ils se distinguent par les spores elliptiques.
Family: Pezizaceae
Pachyphlodes Zobel
in Corda, Icon. fung., 6: 55 (1854)
Type species: Pachyphloeus ligericus Tul. & C. Tul. 1851
Type designated by Zobel, in Corda (1854)
Diagnosis: Episporium muricatum.
Family: Pezizaceae
Pachyphloeus Tul. & C. Tul., nom. illeg.
Giorn. bot. ital., 1 (7-8): 60 (1845)
Type species: Choiromyces melanoxanthus Tul. & C. Tul. ex Berk. 1844
Type designated by Tulasne & Tulasne (1845)
Diagnosis: Stratum corticale peridii vices gerens crassum celluloso-carnosum, extus minute obtuseque verrucosum, apicem versus diminutum aut saepius foramine circulari lato crasse obtuseque marginato rimave tomento peculiari occlusis pervium, nec non et basi absorbenti minutissima non producta suberadicata instructum. Moles interior carnosa molliuscula venis primum marmorata, tandem subuniformis et unicolor; venarum natura duplex, aliae nempe pallidae ab initio coloratae et latiores saepissime et praecipue apicem versus demum hiantes et excavatae, e variis uteri partibus ad ostium vel fungi culmen rimosum confluentes, aerem videlicet admittum vehentes, aliae contra ex inferiore strati corticali facie in plantae infimis enatae, venis alteris intermediis obscuriores, utrinque sporangia gignentes. Sporangia ideo venas steriles vertice spectantia, oblonga lagenaeformia breviter pedicellata simplici plerumque serie ordinata. Sporae octo reticulato-echinatae sphaericae in quoviscunque conceptaculo inordinate nascentes.
Synonym of Pachyphlodes
Paradiscina Benedix
Kulturpflanze, 17: 274 (1969)
Type species: Discina melaleuca Bres. 1898
Type designated by Benedix (1969)
Family: Incertae sedis
Number of species: 1
Current definition: Ascomata epigeous, sessile or substipitate; hymenophore cupulate to discoid, dark brown to black; stipe up to 1 cm long and wide, white, fluted. Ascospores hyaline, ellipsoid, nonapiculate, slightly thickened at ends; surface warty to coarsely rugose; perispore cyanophilic; contents triguttulate.
Paradoxa Mattir.
Beitr. Kryptfl. Schweiz, 8 (2): 32 (1935)
Type species: Paradoxa monospora Mattir. 1935
Family: Tuberaceae
Number of species: 1 fide DoF
Paragalactinia Van Vooren
Ascomycete.org, 12 (4): 188 (2020)
Type species: Peziza succosa Berk. 1841
Type designated by Van Vooren (2020)
Family: Pezizaceae
Current definition: Ascomata hypogeous or epigeous; apothecial species sessile and cupulate. Flesh producing watery latex, becoming yellowsih, greenish or bluish in air. Asci operculate, 8-spored, mainly without crozier, wall weakly bluing in contact with an iodine solution, a little more intensely at the top (WT type). Paraphyses containing a brownish vacuolar pigment. Medullary excipulum containing laticiferous hyphae. Ectal excipulum of textura globulosa, with large cells. Ascospores uni- or biguttulate, warted. Species forming ectomycorrhizas.
Paragyromitra X.C. Wang & W.Y. Zhuang
Microbiol. Spectrum, 11 (3): 10 (2023)
Type species: Helvella infula Schaeff. 1774
Type designated by Wang et al. (2023)
Family: Discinaceae
Number of species: 8
Current definition: Ascomata epigeous, stipitate; hymenophore saddle-shaped, cerebriform, or irregularly lobed, yellow–brown, reddish brown, dark brown, or purple; stipe subcylindric, sometimes with longitudinal furrows, hollow, yellow–brown to light brown or purple. Ascospores hyaline, narrowly ellipsoid, ellipsoid, or fusoid, nonapiculate or infrequently with broadly rounded apiculi; surface smooth to rugulose at maturity; perispore cyanophilic; contents biguttulate.
Paramitra Benedix
Kulturpflanze, 10: 368 (1962)
Type species: Paramitra ochreoides Benedix 1962
Type designated by Benedix (1962)
Synonym of Peziza
Paranthracobia Van Vooren & Valade
Ascomycete.org, 15 (4): 128 (2023)
Type species: Anthracobia rehmii Brumm. 1984
Type designated by Van Vooren & Valade (2023)
Diagnosis: Differs from Anthracobia by the presence of true excipular and marginal hairs, pluriguttulate ascospores and absence of carotenoid pigments
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, slightly cupuliform, disciform to discoid, whitish, pale greyish, orangish or ochre-brown, with an external surface covered by small brown hairs. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura globulosa/angularis. Hairs ± superficial, septate, with a simple base, short, straight. Ascospores uniseriate, ellipsoid or fusoid, hyaline, guttulate, smooth. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored. Paraphyses without carotenoid pigments. Species saprobic, on burnt substrates. Asexual morph unknown.
Parascutellinia Svrček
Česká Mykol., 29: 129 (1975)
Type species: Parascutellinia violacea (Velen.) Svrček 1975
Family: Pyronemataceae
Paratricharina Van Vooren, U. Lindem., M. Vega, Ribes, Illescas & Matočec
Ascomycete.org, 7 (3): 110 (2015)
Type species: Paratricharina poiraultii (Boud.) Van Vooren, U. Lindemann, M. Vega, Ribes, Illescas & Matočec 2015
Family: Pyronemataceae
Current definition: Apothecia about 10–20 mm in diameter or less, sessile, slightly cupulate with inrolled margin, expanded when old; outer surface covered by dense brown-red hairs, sometimes appressed and/or forming distinct pustules, but mostly straight and projecting outwards. Subhymenium thin, of textura intricata. Medullary excipulum of textura intricata, with thin-walled hyphae. Ectal excipulum of vertically oriented textura prismatica, with thin-walled cells, becoming a textura globulosa/subangularis in the outermost part, then with thick-walled brownish cells. Hairs more or less thick-walled, pale brown, more or less sinuous, septate, obtuse or pointed, with a simple base arising from the globose cells of the outer layer of the ectal excipulum. Asci cylindrical, with forked base, arising from perforated croziers, inamyloid, 8-spored. Paraphyses cylindrical, rounded and slightly enlarged at the apex, with spumous compound KOH soluble content, without reaction in Lugol’s solution. Ascospores ellipsoid, hyaline, appearing eguttulate (in dead state) but with small polar aggregations of minute lipid bodies rapidly coalesced into two small polar guttules (in living state), more or less thick-walled, smooth observed in water and Cotton Blue, but with a rough to finely verrucose perispore if mounted in Lugol’s solution, entirely encapsulated by a delicate persistent sheath. Base of apothecia covered by hyaline anchoring hyphae.
Paratrichophaea Trigaux
Docum. Mycol., 16 (61): 5 (1985)
Type species: Paratrichophaea macrocystis Trigaux 1985
Synonym of Lasiobolidium
Parawilcoxina Van Vooren
Ascomycete.org, 13 (1): 36 (2021)
Type species: Parawilcoxina inexpectata Valencia, Van Vooren & M. Vega 2021
Diagnosis: Differs from Wilcoxina by the presence of vacuolar bodies in paraphyses, its saprobic status, and its genetic profile.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, slightly cupuliform, disciform to discoid, whitish to pale greyish, with an external surface covered by small brown hairs. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura globulosa/angularis. Hairs superficial, straight, septate, with a simple base, sometimes arising from a chain of rounded cells. Ascospores uniseriate or irregularly biseriate, ellipsoid to narrow ellipsoid, hyaline, guttulate, smooth. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored. Paraphyses containing vacuolar bodies. Species saprobic. Asexual morph unknown.
Paurocotylis Berk.
Bot. Antarct. Voy. Erebus Terror, 1839-1843 I, Part II, 2: 188 (1855)
Type species: Paurocotylis pila Berk. 1855
Family: Tarzettaceae
Perilachnea Van Vooren
Ascomycete.org, 13 (1): 18 (2021)
Type species: Perilachnea hemisphaerioides (Mouton) Van Vooren 2021
Diagnosis: Differs from Trichophaea s. str. by deeply cupulate apothecia (at least in young state), mainly with a Humaria-like appearance, bi- or pluriguttulate ascospores, paraphyses containing small oil bodies, a saprobic status, and genetic profile.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, deeply cupuliform or discoid, whitish, yellowish or orange, with an external surface covered by sparse brown hairs. Margin hairy. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura globulosa/angularis. Hairs superficial, septate, with a simple base. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored. Paraphyses slender, containing small oil bodies. Ascospores uniseriate, ellipsoid, hyaline, bi- or pluriguttulate, smooth or very finely dotted. Species saprobic of the litter and woody debris of conifers or on enriched or burnt soil. Asexual morph unknown but germinating ascospores observed in P. hemisphaerioides and P. flavobrunnea.
Petchiomyces E. Fisch. & Mattir.
Nat. Pflanzenfam. (Ed. 2), 5bVIII: 40 (1938)
Type species: Petchiomyces thwaitesii (Berk. & Broome) E. Fisch. & Mattir. 1938
Family: Pyronemataceae
Peziza Dill. ex Fr.
Syst. mycol., 2 (1): 40 (1822)
Type species: Peziza vesiculosa Bull. 1790
Type designated by Clements & Shear (1931)
Diagnosis: Fungoides Tournef. J. R. H. p. 564. Fungoid. cava. Mich. gen. p. 205. Peziza. Dill. gen. p. 74. Ellvellae spec. Gled Schaeff. &c. Terana, Pezica, Trombetta. Adons. Octospora, Hedw. Musc. II. p. 4. Patella. Weber. Encoelia Hill. Pezizae spec. Pers. syn. p. 631. Nees Syst. p. 254.
CHAR. Receptaculum marginatum, cupulaeforme, primo sabelausum, mox expansum, epidermide tenui contigua. Hymenium laeve, persistens, distinctum. Asci ampli, distincti, fixi, sporidia elastice ejicienda includentes, parapbysibus immixtis. — Cupula centro adfixa, passim stipitata, libere evoluta, plus minus cava, demum passim planiuscula, disco polito (nec villoso l. pulveraceo) discolori. Velum universale, sed non distinctum & saepe nullum. Substantia carnose-membranacea l. ceracea.
Family: Pezizaceae
Asexual morph of Chromelosporium, Oedocephalum type.
Pfistera Korf & W.Y. Zhuang
Mycotaxon, 40: 275 (1991)
Type species: Pfistera pyrophila Korf & W.Y. Zhuang 1991
Type designated by Korf & Zhuang (1991)
Diagnosis: Ab Pezizalibus aliis (praeter Ascozonus) asci tholo apicale gaudentibus pariete valde tenui munito, ab Ascozono habitu non coprophilo et apotheciis multo magis grandioribus et complexe fabricatis differens.
Synonym of Peziza
Phaeangium Pat.
J. Bot. Paris, 8: 155 (1894)
Type species: Phaeangium lefebvrei Pat. 1894
Synonym of Picoa
Phaeopezia (Vido) Vido
Michelia, 1 (5): 595 (1879)
Type species: Peziza apiculata Cooke 1875
Type designated by Vido (1879)
Diagnosis: Est Geoscypha sporidiis apiculatis fuscisque instructa.
Family: Pezizaceae
Current definition: Ascomata epigeous or hypogeous; apothecial species sessile and discoid. Asci operculate, 8-spored, with crozier, wall diffusely bluing in an iodine solution (WT type). Paraphyses diffusely coloured (olivaceous brown), with also a brownish external pigment on the top article. Ascospores containing numerous small polar granules, ornamented with isolated warts and with polar apiculi, hyaline becoming brown with age. Excipulum of textura globulosa, with small to medium-sized cells. Species forming ectomycorrhizas.
Phillipsia Berk.
J. Linn. Soc. Bot., 18: 388 (1881)
Type species: Phillipsia domingensis Berk. 1881
Family: Sarcoscyphaceae
Phylloscypha Van Vooren
Ascomycete.org, 12 (4): 189 (2020)
Type species: Peziza phyllogena Cooke 1877
Type designated by Van Vooren (2020)
Family: Pezizaceae
Current definition: Ascomata epigeous, cupulate, sessile, external surface distinctly furfuraceous or pustulate. Flesh without latex, purplish-coloured. Asci operculate, 8-spored, with crozier, wall diffusely bluing in an iodine solution (W type). Paraphyses hyaline, with an external pigment at the top, pale brown, olivaceous brown or dark brown. Ascospores eguttulate but containing small polar granules, ornamented with warts. Species saprobic.
Phymatotrichopsis Hennebert
Persoonia, 7 (2): 199 (1973)
Type species: Ozonium omnivorum Shear 1907
Family: Rhizinaceae
Number of species: 1 fide DoF
Physomitra Boud.
Bull. Soc. mycol. Fr., 1: 99 (1885)
Type species: Gyromitra esculenta Pers. ex Fr. 1849
Synonym of Gyromitra
Picoa Vittad.
Monogr. Tuberac.: 54 (1831)
Type species: Picoa juniperi Vittad. 1831
Family: Pyronemataceae
Pindara Velen.
Monogr. Discom. Bohem.: 341 (1934)
Type species: Pindara terrestris Velen. 1934
Type designated by Velenovský (1933)
Diagnosis: Apoth. rigide carnosa, mediocria, crasse stipitata, stipite adulto sulcata, primum leniter patellaria, dein discino-explanata, hymenio colorato, glabra. As. permagni, jodo fusci, par. simpl. filif., apice clavatae. Sp. maximae, distichae, cylindro-fusiformes, hyalinae, serie guttularum magnarum donatae, unicellulares.
Unica species, ad terram humidam ad rivulos in societate Lachnearum aestate nascens. Typus Pezizae eximius, sed meo sensu in vicinitatem Acetabulae spectans. Ab ca apotheciis vestutis discinis, fere pileatis, sporis maximis, cylindricis praeter alia differt.
Family: Helvellaceae
Piscidiscina Dirks, Methven & A.N. Mill.
Mol. Phylogen. Evol., 205: 11 (2025)
Type species: Discina leucoxantha Bres. 1881
Type designated by Dirks et al. (2025)
Family: Discinaceae
Current definition: Ascomata epigeous, substipitate; hymenophore cupulate to discoid, margin incurved; hymenium pinkish-buff to yellow-ochraceous; stipe fluted and lacunose, white. Ascospores hyaline, ellipsoid to fusoid, apiculus truncate with a concave, apical depression; surface roughened to reticulate at maturity; perispore cyanophilic; contents uniguttulate or triguttulate.
Pithya Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 317 (1870)
Type species: Peziza pitya Pers. 1800 (= Pithya vulgaris Fuckel)
Type designated by Seaver (1927)
Diagnosis: Cupulae subsparsae, media magnitudine, ceraceae, primo teretes, demum explanatae, in stipitem crassum brevem attenuatae, basi albicante fibrillosae, ceterum subglabrae, margine acuto vix distincto, disco colorato. Asci longissimi, cylindracei, 8spori. Sporidia monosticha, perfecte globosa, simplicia, hyalina dilutissime rubellave. Paraphyses filiformes, clavatae. Autumnales.
Dürre Aestchen, bisher nur der Coniferen bewohnend.
Family: Sarcoscyphaceae
Asexual morph of Mollardiomyces type.
Planamyces Crous & Decock
Persoonia, 39: 377 (2017)
Type species: Planamyces parisiensis Crous & C. Decock 2017
Family: Otideaceae
Number of species: 1
Plectania Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 323 (1870)
Type species: Peziza melastoma Sowerby 1799
Type designated by Saccardo (1889)
Diagnosis: Cupulae sparsae, majusculae, crasso-stipitatae, carnosae, basi plus minusve comatae, ceterum subglabrae, marginatae, disco plane concavo, discolori. Asci longissimi, cylindracei, 8spori. Sporidia monosticha, oblonga, demum guttulata, continua, episporio laevi, hyalino subrubellove. Paraphyses filiformes, subsimplices. Vernales.
Faulende Aestchen und dergl. bewohnend.
Family: Sarcosomataceae
Asexual morph of Conoplea type.
Plicaria Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 325 (1870)
Type species: Peziza trachycarpa Curr. 1864
Type designated by Boudier (1885)
Diagnosis: Cupulae gregariae sparsaeve, a minoribus ad maximas, carnosae, substipitatae sessilesve, lato-obconicae, turbinatae vel explanato-plicatae, extus aut villo denso, brevi, aut furfuraceae, rarius subglabrae, margine elevato, disco plerumque nitido, concavo explanatove, discolori, quandoque tremelloso. Asci magni, cylindracei, subvermiculares, substipitati, fusci, toti oblique transverse plicati, 8spori. Sporidia in asci superiori parte primo disticha, demum monosticha, perfecte globosa, ovata oblongave, continua, episporio crasso, tuberculoso, fusco (in P. violaceae modo laevi hyalinoque). Paraphyses filiformes, articulatae, demum secedentes (an omnium ?). Terrestres fimicolaeque. Peziza trachycarpa Curr. in Rbh. F. eur. 620. et Peziza Marsupium Pers. Myc. eur. I. pag. 228. etiam hujus generis sunt.
Family: Pezizaceae
Asexual morph of Chromelosporium type.
Plicariella (Sacc.) Rehm
Rabenh. Krypt.-Fl. (ed. 2), 1 (3), lief. 42: 993 (1894)
Type species: Peziza radula Berk. & Broome 1877
Type designated by Eckblad (1968)
Diagnosis: Ascomata subcupulata, majuscula. Sporidia globosa. [P. radula (B. ef Br.) Sacc, P. atrospora (Fuck.) Sacc, P. scabrosa (Cooke) Sacc].
Family: Pezizaceae
Pseudaleuria Lusk
Mycotaxon, 30: 419 (1987)
Type species: Pseudaleuria quinaultiana Lusk 1987
Diagnosis: Apothecia sparsa vel fasciculata, mediocria, sessilia vel substipitata, orbicularia vel repanda; hymenio clare rubro-aurantiaco, extus similari vel pallidiore, pilis hyalinis, flexuosis, stratum externum velutinum tormantibus; excipulo duobus stratis composito, strato interiore ex textura intricata, strato exteriore ex textura angularis, ascis operculatis, non amyloideis, octosporis, cylindraceis, basin versus sensim angustatis, ad maturitatem non prodtrudentibus, muro tenui; ascosporis ellipsoideis, laevibus, hyalinis, eguttulatis, mediocribus; paraphysibus gracilibus, sursum plus minsve dilatatis, raro ramificantibus.
Family: Pyronemataceae
Current definition: Apothecia scattered or in clusters, medium sized, sessile to substipitate, discoid to repand, thick-fleshed, texture firm fleshy to rubbery, becoming corky when dry; hymenium bright reddish orange; exterior concolor or lighter when fresh, with flexuous hyphoid hairs, these hairs forming a felty layer; excipulum two-layered, medullary layer is textura intricata, ectal layer is textura angularis; asci operculate, non-amyloid, eight-spored, cylindrical, tapering to the base, thin-walled, not protruding at maturity; ascospores ellipsoid, smooth, hyaline, eguttulate, uninucleate, medium sized; paraphyses slender, slightly to greatly and abruptly enlarged above, rarely branching.Soil and/or wood saprobe, or mycorrhizal.
Pseudoboubovia U. Lindem., M. Vega, B. Perić & R. Tena
Z. Mykol., 81 (2): 391 (2015)
Type species: Kotlabaea benkertii B. Perić 2013
Family: Pyronemataceae
Current definition: Apothecia 1-8 mm in diameter, sessile, smooth, at first hemispherical, later becoming disc-shaped, finally with irregular shape. Hymenium lemon yellow, greenish yellow, orange yellow. Margin slightly raised, smooth or finely serrated, sometimes irregularly wavy or incurved, same color as the hymenium. Asci 8-spored, operculate, not amyloid, base simple. Ascospores ellipsoid-ovoid or rhomboid, smooth, hyaline or occasionally pale yellowish, with many minute refractive droplets, later becoming thick-walled and droplets accumulating in the centre, with a cyanophilic perispore, dead ascospores containing occasionally a de-Bary-bubble. Paraphyses filiform, distinctly curved, septate, containing yellow or orange granules which do not stain green with iodine. Excipulum two-layered: medullary excipulum of textura intricata, composed of thin-walled hyaline cells; ectal excipulum as textura globulosa, composed of thin walled hyaline cells, intermixed with swollen hyphae. Anchor hyphae covering the base of apothecia.
Pseudocoprotus U. Lindem., Fellm. & J.A. Castillo
Ascomycete.org, 11 (1): 20 (2019)
Type species: Cheilymenia catenipila J. Moravec 2003
Diagnosis: Differs from the species of Coprotus sensu Korf & Kimbrough by its marginal hairs composed of globose and elongate cells in chains, its fully developed marginal tissue, its much bigger size, and the rDNA analysis.
Family: Ascodesmidaceae
Pseudodiscina X.C. Wang & W.Y. Zhuang
Microbiol. Spectrum, 11 (3): 12 (2023)
Type species: Peziza melaleucoides Seaver 1928
Type designated by Wang et al. (2023)
Family: Discinaceae
Current definition: Ascomata epigeous, subsessile to stipitate; hymenophore cupulate to discoid; hymenium rugose, dark gray-brown to brown; stipe short, fluted. Ascospores hyaline, ellipsoid to broadly ellipsoid, nonapiculate; surface with isolated, rounded warts at maturity; perispore cyanophilic; contents biguttulate.
Pseudombrophila Boud.
Bull. Soc. mycol. Fr., 1: 106 (1885)
Type species: Pseudombrophila pedrottii Boud. 1885
Family: Pseudombrophilaceae
Pseudopithyella Seaver
N. Amer. Cup-fungi (Opercul.): 153 (1928)
Type species: Sarcoscypha minuscula Boud. & Torrend 1911
Family: Sarcoscyphaceae
Pseudoplectania Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 324 (1870)
Type species: Peziza nigrella Pers. 1801
Type designated by Boudier (1885)
Diagnosis: Cupulae gregariae sparsaeve, sessiles vel breviter stipitatae, majusculae, carnosae, extus aut pilosae, praecipue basin versus comatae, aut subglabrae, stipite basi in hyphas soluto, effuse radicante, disco marginato, concavo, discolori. Asci longissimi, cylindracei, 8spori. Sporidia monosticha, perfecte globosa, simplicia, laevia, hyalina vel pallide fusca. Paraphyses filiformes, simplices, subclavatae, quandoque coloratae. Vernales.
Auf faulenden Tannennadeln, selten auf nackter Erde.
Verhält sich zu Plectania wie Crouania zu Leucoloma.
Family: Sarcosomataceae
Pseudorhizina Jacz.
Opredelitel’ Gribov., 1. Sovershennye Griby: 414 (1913)
Type species: Pseudorhizina korshinskii Jacz. 1913
Type designated by Jaczewski (1913)
Family: Discinaceae
Current definition: Ascomata epigeous, stipitate; hymenophore irregularly convex, campanulate, saddle-shaped, or irregularly lobed, margin reflexed, fused, or free; hymenium brown to gray-brown or blackish-brown; stipe lacunose, sharply costate, ribs extending onto under surface of hymenophore, white to rosy pink to violet. Ascospores hyaline, globose to ellipsoid, nonapiculate; surface seen as smooth with light microscopy but smooth or finely rugose under SEM at maturity; cyanophilic perispore-periplasm complex absent at maturity; contents uni- or biguttulate. Hyphae of excipulum thick-walled.
Pseudosarcosoma M. Carbone, Agnello & P. Alvarado
Ascomycete.org, 5 (1): 6 (2013)
Type species: Sarcosoma latahense Paden & Tylutki 1969
Type designated by Carbone et al. (2013)
Family: Chorioactidaceae
Current definition: Apothecia turbinate to discoid, gelatinous to less gelatinous in age, substipitate, hymenium deep purple to black, external surface grayish to black. Asci operculate, 8-spored, inamyloid, tapering basally with some forked bases; spores elliptical to subcylindrical, smooth, hyaline, oil-dropped to eguttulate at maturity; paraphyses cylindroid, branched, anastomosing, closely septate, with bent or lobed tips; subhymenium of a thick textura intricata; medullary excipulum of a loose textura intricata; ectal excipulum of textura angularis, the most outer cells with brown walls; external hairs branched, septate, smooth, lobed, hyaline to pale brownish-olivaceous.
Pseudotis (Boud.) Boud.
Hist. Class. Discom. Eur.: 52 (1907)
Type species: Peziza abietina Pers. 1794
Synonym of Otidea
Pseudotricharina Van Vooren, Tello & M. Vega
Ascomycete.org, 11 (2): 32 (2019)
Type species: Pseudotricharina intermedia Van Vooren, Tello & M. Vega 2019
Diagnosis: Differs from the known species of Tricharina by its guttulate and roughly warted ascospores, and from Trichophaea species by its tricharinoid general morphology.
Family: Pyronemataceae
Pseudoverpa (P.A. Moreau, Bellanger & Loizides) X.C. Wang & W.Y. Zhuang
Microbiol. Spectrum, 11 (3): 14 (2023)
Type species: Gyromitra anthracobia Loizides, P.A. Moreau & Bellanger
Type designated by Wang et al. (2023)
Family: Discinaceae
Current definition: Ascomata epigeous, stipitate; hymenophore cerebriform to lobate; hymenium gray-brown, red-brown, purple-brown, or black when fresh; stipe cylindric, attached to hymenophore only at apex, white to ochraceous-orange, stuffed to hollow. Ascospores hyaline, ellipsoid, nonapiculate; surface smooth at maturity; perispore cyanophilic; contents biguttulate.
Psilopezia Berk.
London J. Bot., 6: 325 (1847)
Type species: Psilopezia nummularia Berk. 1847
Family: Rhizinaceae
Ptychoverpa Boud.
Hist. Class. Discom. Eur.: 34 (1907)
Type species: Morchella bohemica Krombh. 1828
Synonym of Verpa
Pulparia P. Karst.
Bidr. Känn. Finl. Nat. Folk., 19: 9 (1871)
Type species: Pulparia arctica P. Karst. 1871
Synonym of Pulvinula
Pulvinula Boud.
Bull. Soc. mycol. Fr., 1: 107 (1885)
Type species: Peziza convexella P. Karst. 1869
Family: Pulvinulaceae
Purpureodiscus (Hirsch) Van Vooren
Ascomycete.org, 12 (4): 189 (2020)
Type species: Galactinia subisabellina Le Gal 1967
Type designated by Hirsch (1992)
Diagnosis: Ascomata minuta vel media, applanata, crassa, estipitata, obscure purpurea vel purpureo brunnea. Carne molle, sine lactae. Sporae hyalinae, laeves, eguttatae. Asci cylindracei, totis parietibus tenuiter amyloidei. Paraphyses simplices, rectes, crassae claviformes, pigmentis in parietibus. Anatomia ascomatis simplex, textura intricata vel inflata, sine mediostrato.
Family: Pezizaceae
Current definition: Ascomata discoid or pulvinate, sessile, fleshy, whitish, pinkish to reddish-colored or purplish brown. Flesh without latex. Asci operculate, 8-spored, with crozier, with wall diffusely bluing in an iodine solution (W type). Paraphyses containing small vacuolar bodies. Ascospores eguttulate, but often with small granules at young stage, smooth or ornamented by isolated warts, in some species germinating to directly produce conidia. Ectal excipulum of textura globulosa/angularis, with large cells. Species saprobic.
Pustularia Fuckel, nom. illeg.
Jahrb. nassau. Ver. Naturk., 23-24: 328 (1870)
Type species: Peziza cupularis L. 1753
Diagnosis: Cupulae sparsae, plerumque maximae, carnosae, breviter crasse stipitatae, campanulatae ampliataeve, extus pustulato-furfuraceae, disco vix discolori, submarginato. Asci longissimi, cylindracei, 8spori. Sporidia in asci superiori parte oblique monosticha, ovata, oblongo-ovatave, continua, 1—2 guttulata, episporio laevi, hyalino, rarius lutescente. Paraphyses filiformes. Terrestres autumnalesque.
Synonym of Tarzetta
Pustulina Eckblad
Nytt Mag. Bot., 15(1-2): 84 (1968)
Type species: Peziza cupularis L. 1753
Type designated by Eckblad (1968)
Synonym of Tarzetta
Pyronema Carus
Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur., 17 (1): 370 (1835)
Type species: Pyronema marianum Carus 1835
Family: Pyronemataceae
Pyropyxis Egger
Can. J. Bot., 62 (4): 705 (1984)
Type species: Peziza rubra Peck 1872
Family: Pyropyxidaceae
Asexual morph of Dichobotrys type.
Ramsbottomia W.D. Buckley
Trans. Br. mycol. Soc., 9 (1): 44 (1924)
Type species: Ramsbottomia lamprosporoidea W.D. Buckley 1924
Family: Pyronemataceae
Reddellomyces Trappe, Castellano & Malajczuk
Aust. Syst. Bot., 5 (5): 606 (1992)
Type species: Labyrinthomyces westraliensis G.W. Beaton & Malajczuk 1986
Diagnosis: A Labyrinthomyces e Dingleya peridio laevi, glabro et ascis sporis 1–4 (–5) differt.
Family: Tuberaceae
Current definition: Differing from Labyrinthomyces and Dingleya in its smooth, glabrous peridium and asci with only 1–4 (–5) spores. Ornamentation of spores of some species collections swells inordinately in KOH. True ornamentation can be viewed in H20, Melzer''s reagent or cotton blue.
Rhizina Fr.
Observ. mycol., 1: 161 (1815)
Type species: Rhizina undulata Fr. 1815
Family: Rhizinaceae
Rhizoblepharia Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 104 (1968)
Type species: Rhizoblepharia jugispora Rifai 1968
Type designated by Rifai (1968)
Family: Pyronemataceae
Number of species: 2 fide DoF
Rhizopodella (Cooke) Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Peziza melastoma Sowerby 1798
Synonym of Plectania
Rhodopeziza Hohmeyer & J. Moravec
Czech Mycol., 47 (4): 261 (1995)
Type species: Aleuria tuberculata Gamundí 1975
Type designated by Moravec (1994)
Diagnosis: Apothecia media usque magna, sessilia, cupuliformia, inaequalia, interdum auriculiformia, externe glabra, ad marginem pruinosa; hymenio miniato. Excipulum externum textura globulosa angularis compositum e cellulis latioribus, excipulum internum (medulla) textura globulosa angularis compositum e cellulis angustioribus cum hyphis intermixtis. Asci cylindracei, operculati, octospori, in toto iodo coerulescentes. Ascosporae late ellipsoideae, eguttulatae, pallide flavescentes, tuberculatae. Paraphyses filiformes, simplices, septatae, rectae sed apice curvatae, granulis aurantiacis impletae.
Family: Pezizaceae
Rhodoscypha Dissing & Sivertsen
Mycotaxon, 16 (2): 442 (1983)
Type species: Peziza ovilla Peck 1876
Family: Pyronemataceae
Rhodotarzetta Dissing & Sivertsen
Mycotaxon, 16 (2): 453 (1983)
Type species: Pustularia rosea Rea 1924
Family: Pyronemataceae
Rickiella Syd. & P. Syd. ex Rick
Ann. mycol., 2 (3): 244 (1904)
Type species: Rickiella transiens Syd. & P. Syd. ex Rick 1904
Family: Sarcoscyphaceae
Rubelia Nieuwl.
Am. Midl. Nat., 4: 386 (1916)
Type species: Peziza trechispora Berk. & Broome 1846
Diagnosis: nomen novum, based on Sphaerospora (Vido) Sacc., illeg.
Synonym of Scutellinia
Rugosporella Pfister, Healy & LoBuglio
Ascomycete.org, 16 (3): 188 (2024)
Type species: Peziza atrovinosa Cooke 1875
Type designated by Pfister et al. (2024)
Family: Pezizaceae
Current definition: Apothecia shallow cupulate, often in groups, hymenium ranging in color from tawny buff to dark brown to nearly black on drying, outer surface concolorous or lighter than the hymenium, scurfy. Flesh thick, composed of textura intricata with intermixed globose cells. Outer excipulum with low pustules composed of parallel cells oriented perpendicularly to the outer surface. Ascospores broadly ellipsoid, under 15 µm long, with two guttules, surface ornamented with a regular or irregular reticulum the veins of which become brown at maturity. Asci blue in iodine at the apex without a ring and blue in the upper half. Paraphyses slightly swollen at the tip, containing dark amorphous material.
Ruhlandiella Henn.
Hedwigia, 42 (Beibl.): 24 (1903)
Type species: Ruhlandiella berolinensis Henn. 1903
Family: Pezizaceae
Current definition: Fruitbodies pulvinate to brain-like. Excipulum of globose to angular, thick-walled cells. Asci cylindrical to clavate, weakly amyloid all over, without operculum; ascus wall evanescent. Paraphyses septate, branching, with a conspicuous gelatinous sheath; paraphyses distinctly exceeding asci in length. Ascospores globose, at first bi- to tri-seriate, hyaline, later uniseriate, with a pale brownish, strongly cyanophilic ornamentation.
Saccobolus Boud.
Ann. Sci. Nat. Bot., sér. 5, 10: 228 (1869)
Type species: Ascobolus kervernii P. Crouan & H. Crouan 1858
Type designated by Boudier (1885)
Diagnosis: Ascobolus, Pers. et omnium auctorum fecentiorum.
Receptaculum carnosum, sessile, extus glabrum, nitidum. Hymenium planum dein convexum, lucidum, thecis minus exsertis granula nigra nitida simulantibus vestitum. Paraphyses numerosæ, minus graciles, ad apicem incrassatæ, septatæ, simplices aut ramosæ; intus vix granulosæ, longitudine thecas æquantes, gelatina vix conspicua immersæ. Thecæ curtæ, amplæ, ad apicem subquadratæ, ad basin attenuatæ et tunc subcuneiformes, operculo sæpius triangulari non umbonalo dehiscentes, sporas 8 includentes. Sporæ episporio ceraceo, leve, rarissime minute etl sæpius transversim rimoso, membrana communi inclusæ, maturæ ad extremitatem thecæ varie aggregatæ.
Species minores stercorariæ.
Family: Ascobolaceae
Current definition: Ascomata paragymnohymenial or eugymnohymenial; superficial, sessile; small 0.1-1.0 mm, rarely up to 2.0 mm diameter. Receptacle at first often subglobular or semiglobular but soon pulvinate or lenticular; surface smooth or sometimes covered with spreading bunches of flexuous hyphae; without a distinct margin. Asci shortly clavate, with clearly truncate apex; at least at maturity the tips strongly protruding above the hymenium; the wall staining blue with iodine. Ascospores firmly united in a cluster or package and ejected together, cemented in a more or less regular pattern (Fig. 2) by the episporial pigment or rarely at first loosely united according to such a pattern and finally free or partly free. Paraphyses slender or rather thick, cylindrical or cylindric-clavate; relatively short; sometimes embedded in colourless mucus; contents colourless or yellow. Sometimes intercellular, amorphous, violet or brown pigment between the tips of paraphyses or in the outer layers of the excipulum.
Fimicolous, more rarely on rotten textiles or on substrata dirtied with dung.
Sarcopeziza Loizides, Agnello & P. Alvarado
Ascomycete.org, 10 (4): 179 (2018)
Type species: Peziza sicula Inzenga 1869
Type designated by Agnello et al. (2018)
Family: Pezizaceae
Current definition: Ascomata hypogeous or semi-hypogeous, medium- to large-sized, sarcosphaeroid in aspect and distinctly or indistinctly stipitate, at first angiocarpic, hollow, subglobose to pyriform, with a small apical orifice, gradually tearing concentrically to form erect or arched lacinias; hymenial surface smooth, purplish-violet; asci with fully functioning opercula, , pleurorhynchous, diffusely amyloid over the entire length, more intensely at the apex; paraphyses multiseptate, cylindrical or sometimes moniliform; ascospores monoseriate, ellipsoid, smooth or finely low warted, hyaline or subhyaline and mono- or multiguttulate,; in xeric scrub, dry grasslands and thermomediterranean matorral.
Sarcoscypha (Fr.) Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Peziza coccinea Jacq. 1774
Family: Sarcoscyphaceae
Asexual morph of Mollardiomyces type.
Sarcosoma Casp.
Rabenh. Krypt.-Fl. Edn, 2, 1.3: 497 (1891)
Type species: Sarcosoma globosum (Schmidel) Casp. 1891
Family: Sarcosomataceae
Asexual morph of Verticicladium type.
Sarcosphaera Auersw.
Hedwigia, 8: 82 (1869)
Type species: Peziza macrocalyx Riess 1852
Type designated by Auerswald (1869)
Diagnosis: Fungus suglobosus, undique clausus, deinde Geastri instar irregulariter rumpens, carnosus, extus membranaceo- corticatus, intus strato crasso merenchymatoso; hymenium tenue ex ascis tubulosis 8-sporis paraphysibusque filiformibus ramosis, septatis consistens.
Family: Pezizaceae
Scabropezia Dissing & Pfister
Nord. J. Bot., 1 (1): 102 (1981)
Type species: Peziza scabrosa Cooke 1877
Type designated by Dissing & Pfister (1981)
Diagnosis: Apothecia raro plus quam 2 cm lata, catilliformia vel cupulata, hymeniis viridulis flavido vel olivaceo affectis, extra pustulis fuscidis prominentibus. Excipulum exterius textura globulosa vel angulari, cellulis globularibus vel elongatis, parietibus fuscidis, extrinsecus in pustulas conspicuas pyramidales vel conicas elevatum; excipulum medullare textura intricata. Asci omnino amyloides, operculati, cylindrici, basibus latis pleurorhynchis. Paraphyses rectae, septatae, supra matrice colorata granulari circumdatae. Ascosporae globulares, initio achroae, in duas series vel sine ordine dispositae, postea fuscidae, verrucosae uniseriales. Habitat in solo et in frustulis ligneis, locis deustis non affinis.
Species typifica: Peziza scabrosa Cooke.
Synonym of Plicariella
Scodellina Gray
Nat. arrang. Brit. plants, 1: 668 (1821)
Type species: Peziza vesiculosa Bull. 1790
Type designated by Eckblad (1968)
Diagnosis: Thallus fleshy, membranaceous, brittle, sessile, hemispherical, spreading; outside scaly or mealy.
Synonym of Peziza
Scotopezia Van Vooren, Sammut & P. Alvarado
Cryptog. Mycol., 44 (12): 166 (2023)
Type species: Peziza pseudoanthracina Donadini 1982
Type designated by Sammut et al. (2023)
Family: Pezizaceae
Number of species: 1
Current definition: Ascomata gregarious, epigeous, discoid, sessile, brown to black coloured. Flesh without latex. Asci operculate, with croziers, having a diffusely but inconstantly amyloid wall in iodine solution. Ascospores globose, uniguttulate, warted. Paraphyses embedded at the top with a yellowish-brown exudate. Excipulum of textura globulosa/subglobulosa. Anamorph unknown. Trophic status unknown.
Scutellinia (Cooke) Lambotte
Mém. Soc. roy. Sci. Liège, sér. 2, 14: 299 (1887)
Type species: Peziza scutellata L. 1753
Family: Pyronemataceae
Selenaspora R. Heim & Le Gal
Rev. Mycol. Paris, 1: 307 (1936)
Type species: Selenaspora batava R. Heim & Le Gal 1936
Type designated by Heim & Le Gal (1936)
Family: Pyronemataceae
Sepultaria (Cooke) Boud.
Bull. Soc. mycol. Fr., 1: 104 (1885)
Type species: Peziza sepulta Fr. 1851
Type designated by Boudier (1885)
Diagnosis: Cupula carnosa, sessilis, extus pilis coloratis vestita.
Family: Pyronemataceae
Sepultariella Van Vooren, U. Lindem. & Healy
Ascomycete.org, 9 (4): 118 (2017)
Type species: Peziza patavina Cooke & Sacc. 1877
Type designated by Van Vooren et al. (2017)
Diagnosis: Differs from Leucoscypha Boud. by pigmented apothecia and smooth ascospores.
Family: Pyronemataceae
Smardaea Svrček
Česká Mykol., 23: 90 (1969)
Type species: Ascobolus amethystinus W. Phillips 1875
Type designated by Svrček (1969)
Diagnosis: Apothecia solitaria usque gregaria, intermedia, patellaria, sessilia, carnosa, tota violacea, extus nuda, disco concolore absque ascis prominulis. Excipulum cum textura globulosa e cellulis globosis vel crasse cylindraceis, violaceo-coloratis constat. Medulla cum textura intricata e hyphis longis, angustis, pallide violaceis constat. Asci cylindracei, octospori, membrana violaceo-colorata, non amyloidea. Paraphyses crassae, septatae, apice rectae, violaceo-coloratae. Sporae oblongo-ellipsoideae, verrucosae, maturae violaceae usque fuscoviolaceae.
Hab. Ad terram humidam.
Family: Pyropyxidaceae
Smarodsia Raitv. & Vimba
Folia cryptog. Estonica, 42: 98 (2006)
Type species: Smarodsia stollii Raitv. & Vimba 2006
Diagnosis: Genus pyronematacearum Kotlabaea Svrcek similis sed in sporis distincte guttulatis et pilis setiformibus crassiter tunicatis differt.
Family: Pyronemataceae
Current definition: Apothecia terrestrial, minute, broadly sessile, discoid to saucer-shaped, disc pale yellowish to pale ochraceous or orange-yellowish, receptacle concolorous with the disc, externally smooth. Ectal excipulum of hyaline, large-celled textura globulosa. The cells of outer layers have thick, refracting walls. Hairs scanty, hyaline, aseptate, firm-walled to thick-walled, cylindrical to conical, obtusely rounded, rarely almost pointed, straight to slightly flexuous, arising superficially from a basal cell. Asci pleurorhynchous, cylindrical, apically not bluing in MLZ, at first 8-spored but 4-spored at maturity in type species. Ascospores ellipsoid-fusoid, containing 2 big, more rarely 1 big or 1 big and several small lipid globules, sometimes with numerous small globules or granulose cytoplasma, spore wall without changes when heated in lactic acid. Paraphyses apically clavate, fi lled with numerous small yellowish drops in CB.
Sowerbyella Nannf.
Svensk bot. Tidskr., 32: 118 (1938)
Type species: Peziza radiculata Sowerby 1797
Family: Pyronemataceae
Spaniodiscus Van Vooren, U. Lindemann & Roffler
Ascomycete.org, 14 (2): 43 (2022)
Type species: Parascutellinia fuckelii Donadini & Svrček 1985
Type designated by Lindemann et al. (2022)
Diagnosis: Differs from Scutellinia and other genera having an orange hymenium by the combination of the following characters: superficial and simple-based hairs, always smooth thick-walled ascospores, filled with oil droplets, and its genetic profile.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, orange-coloured, with an external surface and margin covered by short brown hairs. Excipulum two-layered: medullary layer of textura intricata, ectal layer of textura globulosa/angularis. Hairs simple, superficial, septate, with a simple base. Asci operculate, arising from croziers, inamyloid, 8-spored. Paraphyses slender, enlarged at the top, containing carotenoid pigments. Ascospores uniseriate, ellipsoid, hyaline, pluriguttulate in living state, smooth. Trophic status probably saprobic. Asexual morph unknown.
Sphaeridiobolus Boud.
Bull. Soc. mycol. Fr., 1: 108 (1885)
Type species: Ascobolus hyperboreus P. Karst. 1871
Synonym of Ascobolus
Sphaerosoma Klotzsch
Fl. Regn. Boruss., 7: 467 (1839)
Type species: Sphaerosoma fuscescens Klotzsch 1839
Family: Incertae sedis
Number of species: 3 fide DoF
Sphaerospora (Vido) Sacc., nom. illeg.
Syll. fung., 8: 188 (1889)
Type species: Peziza trechispora Berk. & Broome 1846
Type designated by Seaver (1927)
Synonym of Scutellinia
Sphaerosporella (Svrček) Svrček & Kubička
Česká Mykol., 15 (2): 66 (1961)
Type species: Peziza brunnea Alb. & Schwein. 1805
Synonym of Trichophaea
Sphaerozone Zobel
in Corda, Icon. fung. (Prague), 6: 53 (1854)
Type species: Sphaerozone tulasnei Zobel 1854
Type designated by Zobel, in Corda (1854)
Diagnosis: Fungus cavus, irregulariter gibberoso-plicatus, apice hians, basi mycelio copioso pallide fusco donatus. Peridium tenuissimum (evanidum?). Hymenium periphericum, superficiale, componitur ex ascis clavato-cylindricis exotropis, intermixtis paraphysibus multo longioribus filiformi-clavatis. Asci paralleli, approximati 5—6-, rarius octospori. Sporae sphaericae, in ascis uniseriati. Episporium verrucosum, verrucis crassis obtusiusculis.
Family: Pezizaceae
Number of species: 4 fide DoF
Spooneromyces T. Schumach. & J. Moravec
Nord. J. Bot., 9 (4): 426 (1989)
Type species: Spooneromyces laeticolor (P. Karst.) T. Schumach. & J. Moravec 1989
Family: Pyronemataceae
Stephensia Tul. & C. Tul.
C. r. hebd. Séanc. Acad. Sci. Paris, 21: 1433 (1845)
Type species: Stephensia bombycina (Vittad.) Tul. 1851
Synonym of Hydnocystis
Strobiloscypha N.S. Weber & Denison
Mycotaxon, 54: 130 (1995)
Type species: Strobiloscypha keliae N.S. Weber & Denison 1995
Diagnosis: Apothecia sessilia, juventute aliquantum turbinata, maturitate vade crateriformia vel plana, 1.5-5.0(-7.0) mm in diam, succulenta. Hymenium olivaceo-brunneolum vel olivaceo-cinereum. Pagina abhymenialis brunneola vel brunnea, glabra, margo minute pustullata. Sporae ellipsoideae, (18.7-)21-23.7 x (11.8-)12.5-13(-15.1) µm, hyalinae, maturitatae agullulatae, verruculosae, in ascis uniseriate. Asci J-, operculati, 8-sporae, 316-368 x 14.3-18.4 µm. Paraphyses rectae, anguste clavatae, septatae. Subhymenium e excipulum medullosum texturis intricatis, hyphae 5.3-10.5(-15.6) µm in diam. Excipulum ectale texturis globulosis/angularis. In strobilis Cupressacearum e Taxodiacearum.
Family: Strobiloscyphaceae
Tarzetta (Cooke) Lambotte
Mém. Soc. roy. Sci. Liège, sér. 2, 14: 325 (1888)
Type species: Peziza tarzetta Cooke 1877 (= Peziza catinus Holmsk. 1799)
Type designated by Rogers et al. (1971)
Diagnosis: Cupula subintegra, nuda, stipitata; stipite tenui, vel abbreviato, erecto vel radicato.
Family: Tarzettaceae
Temperantia K. Hansen, Healy & Kovács
Mycologia, 103 (4): 837 (2011)
Type species: Mattirolomyces tiffanyae Healy 2003
Type designated by Kovács et al. (2011)
Diagnosis: Peridium constans ex textura intricata. Gleba a peridio non separabilis, solida, marsupiis fertilibus et venis in sicco cremicoloribus. Asci sporis 1–4, paraphyses carentes. Sporae hyalinae, uniguttulatae, globosae, e verrucato porcatae. A Hydnobolite ac Mattirolomycete ordinibus DNA differt.
Family: Pezizaceae
Current definition: Ascoma a stereothecium without a basal mycelial tuft, not changing color with handling or damage, darkening slightly to cream-colored when dry. Excipulum thin, of slightly inflated, interwoven hyphae similar to those of the sterile glebal veins. Gleba solid, lacking paraphyses or channels. Asci 1–4-spored, irregularly arranged in fertile pockets. Spores globose, with one guttule, hyaline and ornamented with blunt-tipped warts and irregularly thickened ridges (Fig. 3j).
Terfezia (Tul. & C. Tul.) Tul. & C. Tul.
Fungi hypog.: 172 (1851)
Type species: Tuber arenarium Moris 1829
Type designated by Trappe (1971)
Diagnosis: Materie fertili in glebulas pulposas aggregata; sporangiis globosis ecaudatis in ipsa absque ordine nididantibus.
Family: Pezizaceae
Terracavicola A.C. Grupe, Kraisit., Guevara & M.E. Sm.
Ascomycete.org, 11 (2): 41 (2019)
Type species: Terracavicola echinospora A.C. Grupe, Kraisit., Guevara & M.E. Sm. 2019
Diagnosis: A hypogeous, creamy-tan to light brown truffle-like ascoma with a roughened peridium that is covered with copious brown hairs (setae), the presence of a weak epithecium which covers the hymenium and prevents active discharge of the ascospores, and unique ITS and 28S DNA sequences.
Family: Pyronemataceae
Thecotheus Boud.
Ann. Sci. Nat. Bot., sér. 5, 10: 235 (1869)
Type species: Thecotheus pelletieri (P. Crouan & H. Crouan) Boud. 1869
Family: Ascobolaceae
Thindia Korf & Waraitch
Mycologia, 63 (1): 102 (1971)
Type species: Thindia cupressi Korf & Waraitch 1971
Family: Sarcoscyphaceae
Tirmania Chatin
Bull. Soc. bot. Fr., 38: 62 (1891)
Type species: Tuber niveum Desf. 1799
Type designated by Trappe (1971)
Family: Pezizaceae
Trichaleurina Rehm
Ann. mycol., 1 (6): 515 (1903)
Type species: Trichaleurina polytricha Rehm 1914
Family: Chorioactidaceae
Tricharia Boud., nom. illeg.
Bull. Soc. mycol. Fr., 1: 104 (1885)
Type species: Tricharia gilva (Cooke) Boud. 1904
Synonym of Tricharina
Tricharina Eckblad
Nytt Mag. Bot., 15 (1-2): 60 (1968)
Type species: Tricharina gilva (Boud. ex Cooke) Eckblad 1968
Family: Pyronemataceae
Asexual morph of Ascorhizoctonia type.
Tricharinopsis U. Lindem., Van Vooren & Healy
Ascomycete.org, 11 (5): 155 (2019)
Type species: Tricharinopsis herinkii (Svrček) U. Lindem., Van Vooren & Healy 2019
Diagnosis: Differs from Tricharina Eckblad by the absence of clustered hairs at the margin and by an outer surface densely covered with long hyaline to brown flexuous hairs (reminiscent of a Geopora species).
Family: Pyronemataceae
Trichobolus (Sacc.) Kimbr. & Cain
Amer. J. Bot., 54 (1): 20 (1967)
Type species: Trichobolus zukalii (Heimerl) Kimbr. 1967
Family: Ascodesmidaceae
Trichophaea Boud.
Bull. Soc. mycol. Fr., 1: 105 (1885)
Type species: Trichophaea woolhopeia (Cooke & W. Phillips) Arnould 1893
Diagnosis: Poils du réceptacle longs et aigus ; spores le plus souvent garnies de gouttelettes intérieures. Hyménium blanchâtre ou glauque.
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, subglobose, discoid or cupuliform, hymenium whitish, pale grey to grey-brown, brown or blackish, with an external surface covered by superficial brown hairs. Margin hairy. Excipulum two-layered: medullary layer thick, of textura intricata and ectal layer of textura subglobulosa/angularis. Hairs superficial, septate, with a simple base. Asci operculate, arising from croziers, inamyloid, 8-spored. Paraphyses slender, equal to enlarged (sometimes capitate) at the top, hyaline. Ascospores uniseriate, ellipsoid or spherical, hyaline, uniguttulate, more rarely biguttulate, thick-walled, smooth. Species forming ectomycorrhizae.
Asexual morph of Dichobotrys type.
Trichophaeopsis Korf & Erb
Phytologia, 24 (1): 18 (1972)
Type species: Trichophaeopsis bicuspis (Boud.) Korf & Erb 1972
Family: Pyronemataceae
Current definition: Ascomata epigeous, apothecial, sessile, pale greyish, with an external surface and margin covered by upward-pointing brown hairs. Excipulum three-layered: medullary layer of textura intricata, ectal layer of textura globulosa/angularis and the outermost part of ectal excipulum of textura prismatica. Hairs simple or bifurcate, deeply inserted in the excipulum, septate, mainly with a simple base. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored, more rarely 4-spored. Paraphyses slender, not enlarged at the top. Ascospores uniseriate, ellipsoid, hyaline, eguttulate, but sometimes containing small granules that can merge into two oil drops in overmature or dead ascospores, smooth or verrucose. Species saprobic, on soil, woody debris or organic matter. Asexual morph unknown.
Tuber P. Micheli ex F.H. Wigg.
Prim. fl. holsat.: 1-112 (1780)
Type species: Tuber aestivum Vittad. 1831
Family: Tuberaceae
Tubipeda Falck
Mykol. Untersuch. Ber., 1 (3): 377 (1923)
Type species: Tubipeda elastica (Bull.) Falck 1923
Synonym of Helvella
Ulurua Trappe, Claridge & Kovács
Mycol. Progr., 9 (1): 140 (2010)
Type species: Ulurua nonparaphysata Trappe & Claridge 2010
Type designated by Trappe et al. (2010)
Diagnosis: Ascomata usque ad 4 cm lata, sulcata, globosa vel turbinata, tomentosa, brunnea. Gleba venis labyrinthinis, vacuis vel hymeniis opponentibus repletis. Sporae hyalinae, late ellipsoideae vel subglobosae, 12–15 x 10–12 μm, verrucis minutis ornatae. Asci in hymenio innati, amyloidei, sporis octo. Paraphyses destitutae. Peridium glebaque cellulis multo inflatis.
Family: Pezizaceae
Number of species: 1
Underwoodia Peck
Ann. Rep. N.Y. St. Mus., 43: 32 (1890)
Type species: Underwoodia columnaris Peck 1890
Family: Helvellaceae
Urceolaria Bonord., nom. illeg.
Handb. Allgem. mykol.: 203, 311 (1851)
Type species: Urceolaria agelaea (Ach.) Ach. 1803
Family: Incertae sedis
Number of species: 1 fide DoF
Urinopezia Van Vooren
Ascomycete.org, 15 (2): 77 (2023)
Type species: Peziza tarembergensis Moyne, Stöckli, C. Page & Van Vooren 2017
Type designated by Van Vooren (2023)
Family: Pezizaceae
Number of species: 1
Current definition: Ascomata epigeous, cupulate. Flesh without latex. Asci operculate, 8-spored, with croziers, wall diffusely bluing in an iodine solution, more intensely at the top (WT type after the typology proposed by Van Vooren, 2020). Paraphyses containing small refractive vacuoles. Medullary excipulum of textura globulosa with large-sized cells, ectal excipulum of textura globulosa with smaller cells. Ascospores guttulate, smooth. Asexual morph unknown. Species saprobic on substrates enriched by urine.
Urnula Fr.
Summa veg. Scand., section post.: 364 (1849)
Type species: Urnula craterium (Schwein.) Fr. 1851
Family: Sarcosomataceae
Velenovskya Albanese, Boragine, M. Carbone & P. Alvarado
Ascomycete.org, 14 (3): 111 (2022)
Type species: Plicariella vacini Velen. 1939
Type designated by Albanese et al. (2022)
Family: Pezizaceae
Current definition: Apothecia discoid or irregular in outline, flattened to more or less convex, smooth to cerebriform; ascospores initially hyaline then yellow-brownish at maturity, ornamented; asci operculate, lightly amyloid in almost their entire length, much deeper at the top, 8-spored, with croziers; paraphyses cylindrical, septate, bifurcated, rarely trifurcated, with apex of various shape immersed in a light-brownish gelatinous content; subhymenium thin, of a dense textura intricata; medullary excipulum of textura intricata of cylindrical, subhyaline hyphae mixed with pyriform to rounded cells in the outer part; ectal excipulum of textura angularis. Besides its unique genetic profile, it differs from Plicariella in having ellipsoid ascospores, and from Peziza s.str. in having heavily ornamented light-brownish ascospores at maturity.
Verpa Sw.
K. svenska Vetensk-Akad. Handl., 36: 129 (1815)
Type species: Verpa conica (O.F. Müll.) Sw. 1815
Family: Morchellaceae
Verticicladium Preuss
Linnaea, 24: 127 (1851)
Type species: Verticicladium trifidum Preuss 1851
Family: Incertae sedis
Number of species: 1 fide DoF
Warcupia Paden & J.V. Cameron
Can. J. Bot., 50 (5): 999 (1972)
Type species: Warcupia terrestris Paden & J.V. Cameron 1972
Family: Otideaceae
Wenyingia Zheng Wang & Pfister
Mycotaxon, 79: 397 (2001)
Type species: Wenyingia sichuanensis Zheng Wang & Pfister 2001
Diagnosis: Apothecia cupulata, stipitata, brunneloa, usque lO mm diam, 22 mm altitudo. Excipulum ectal e cellulis brunneloa, angularibus 5-20 µm diam (textura angulari), excipulum medullare ex hyphis brunneae. Compositum. Asci octospori, crasse tunicati, cylindracei, 270-295 × 13.5-16 µm. Ascospori ellipsoidei, biguttulati, 19-21.5 × 11-12 µm.
Family: Tarzettaceae
Number of species: 1 fide DoF
Wilcoxina Chin S. Yang & Korf
Mycotaxon, 24: 511 (1985)
Type species: Wilcoxina mikolae (Chin S. Yang & H.E. Wilcox) Chin S. Yang & Korf 1985
Diagnosis: Ab Tricharina excipulo ectali non distincte bistrato, ascosporis dimidium superius asci maturi tantum occupantibus, et procreatione anamorphosis lente crescentis generis Complexipedis et non celeriter crescentis generis Ascorhizoctoniae distinguenda.
Family: Pyronemataceae
Current definition: Apothecia small, cupulate, expanding to discoid, sessile, attached only at the center, scattered or gregarious; hymenium whitish to yellowish orange, senescing to brownish gray; hairs both marginal and running down the flanks, fasciculate, with large clusters near margin and clusters gradually becoming smaller towards the base of the apothecium, pale brown to brown, with narrow bases arising from a chain of thick-walled excipular cells, the broadest part of the hair well above the point of origin of the hair, containing brownish, globose inclusions often appearing attached to the walls, walls 2.0-4.0 µm thick, with thin or thick septa; excipulum clearly two-layered near the margin, ental layer becoming broader toward the apothecial base, of textura intricata, in some taxa having refractive, thick-walled hyphae threading horizontally through the layer, ectal layer of textura globulosa to textura angularis, composed of some almost globose to radially elongated cells with pale brown to brown, thick walls becoming mixed near the apothecial base with some small-diameter hyphae from the ental layer; asci cylindrical, narrowing toward base, apically operculate, 8-spored or rarely (2-) 4 (-6)-spored, J-; ascospores ellipsoid, uniseriate, lining up in the upper half of the ascus, smooth, hyaline or refractive-yellow when mature and mounted in media containing lactic acid, eguttulate, but often with a few small granules or with small, polar clusters of fine granules; paraphyses slender, septate, branched near base, with or without carotenoid particles, slightly enlarged toward apex.
Asexual morph of Complexipes type.
Wolfina Seaver ex Eckblad
Nytt Mag. Bot., 15 (1-2): 126 (1968)
Type species: Wolfina aurantiopsis (Ellis) Seaver 1937
Family: Chorioactidaceae
Wynnea Berk. & M.A. Curtis
J. Linn. Soc. Bot., 9: 424 (1867)
Type species: Wynnea gigantea Berk. & M.A. Curtis 1867
Family: Wynneaceae
Wynnella Boud.
Bull. Soc. mycol. Fr., 1: 102 (1885)
Type species: Wynnella auricula (Schaeff.) Boud. 1885
Type designated by Eckblad (1968)
Synonym of Midotis
Last Update on 09/05/2025