Genera of Pezizales

The genera related to Pezizales are listed below. Those in bold are currently accepted. The number of known species is given based on our knowledge or on the Dictionary of Fungi (10th ed., 2008), abreviated in DoF.

A B C D E F G H I J K L M N O P R S T U V W 

 

Acervus Kanouse
Pap. Mich. Acad. Sci., 23: 149 (1938)
Type species: Acervus aurantiacus Kanouse 1938
Family: Otideaceae
Number of species: 4 fide DoF [6 integrated in 2000 Pezizales]

Acetabula (Fr.) Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 330 (1870)
Type species: Acetabula vulgaris Fuckel 1870
Synonym of Helvella

Ahmadea Aman, Khalid & Moncalvo
Stud. Fung., 5 (1): 454 (2020)
Type species: Ahmadea dalanensis Aman & Khalid 2020
Diagnosis: Hypogeous ascomata, irregular in shape, thin excipulum, solid gleba lacking paraphyses, asci with eight spores (sometimes less), ascospores globose, ornamented with sharp spines with a broad base and blunt ends.
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Aleuria Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 325 (1870)
Type species: Aleuria aurantia (Pers.) Fuckel 1870
Family: Pyronemataceae
Number of species: 17 fide DoF [6 integrated in 2000 Pezizales]

Aleurina Massee
Bull. Misc. Inf. Kew, 1898: 13 (1898)
Type species: Aleurina tasmanica Massee 1898
Family: Pyronemataceae
Number of species: 11 fide DoF [5 integrated in 2000 Pezizales]

Amylascus Trappe
Trans. Br. mycol. Soc., 57 (1): 89 (1971)
Type species: Amylascus herbertianus (J.W. Cribb) Trappe 1971
Family: Pezizaceae
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Anthracobia Boud.
Bull. Soc. mycol. Fr., 1: 106 (1885)
Type species: Peziza melaloma Alb. & Schwein. 1805
Family: Pyronemataceae
Number of species: 15 fide DoF [5 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, first cupuliform then discoid or flattened, hymenium yellow-orange to dull orangish, with an external surface covered by superficial light umber brown to brown pseudo-hairs, i.e. brown clavate or obtuse elongated cells. Margin hairy. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura subglobulosa/angularis. Asci operculate, arising from croziers, inamyloid, 8-spored. Paraphyses slender, enlarged to capitate at the top, hyaline. Ascospores uniseriate, ellipsoid, hyaline, biguttulate, thick-walled, smooth. Species saprobic on burnt soil.
Asexual morph of Scytalidium type.

Antrelloides P.S. Catches. & D.E.A. Catches.
Swainsona, 31 (6): 82 (2018)
Type species: Antrelloides atroceracea P.S. Catches. & D.E.A. Catches. 2018
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Apothecial ascomata up to 40 mm diam. Hymenium convex; firm, waxy; black, violaceous-black. Basal stipe substantial; obconic; a chambered base of knobby columnar ridges radiating from base; fragile, waxy. Asci amyloid; cylindrical; operculate; 8-spored. Ascospores ellipsoid; smooth under light microscope. Paraphyses longer than asci; septate; tips swollen and tending to aggregate, encrusted with brown amorphous matter; branching occasionally near tips. Medullary excipulum of several layers of interwoven hyphae, grading into ectal excipulum. Ectal excipulum of large globose to subglobose cells with interwoven hyphae. Stipe tissue of textura globulosa and chains of subglobose cells.

Aparaphysaria Speg.
Anal. Soc. cient. argent., 94 (1-2): 81 (1922)
Type species: Aparaphysaria doelloi Speg. 1922
Family: Pyronemataceae
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Aquapeziza D.M. Hu, L. Cai & K.D. Hyde
Mycologia, 104 (2): 541 (2012)
Type species: Aquapeziza globispora D.M. Hu, L. Cai & K.D. Hyde 2012
Family: Pezizaceae
Number of species: 1 [0 integrated in 2000 Pezizales]
Current definition: Apothecia scattered, epigeous, sessile, turbinate, white; excipulum composed of an ectal exipulum of textura globulosa to textura angularis and medullary excipulum of textura intricata. Paraphyses cylindrical, septate, hyaline. Asci unitunicate, not protruding, operculum not distinct, at first broad cylindrical becoming ovoid at maturity, with a rounded apex, hyaline, amyloid, eight-ascospored. Ascospores globose, one-celled, guttulate, hyaline, smooth.

Arpinia Berthet
Bull. mens. Soc. linn. Lyon, 43, num. spéc.: 36 (1974)
Type species: Arpinia inops Berthet 1974
Diagnosis: Apotheciae terrestres, pedunculatae, cupulares, extrinsecus subtiliter furfuraceae, omnino pallidae, pigmentis carotenoidibus indigentes. Asci octospori, non iodo caerulescentes. Sporae uni – aut biguttulatae, ovatae, laeves, uninucleatae. Paraphyses clavatae, apice rectae. Ascogenae hyphae fibulatae.
Family: Otideaceae
Number of species: 4 fide DoF [3 integrated in 2000 Pezizales]

Ascobolus Pers.
Syst. Nat. Edn, 13, 2 (2): 1461 (1792)
Type species: Ascobolus pezizoides Pers. 1797
Family: Ascobolaceae
Number of species: 61 fide DoF [26 integrated in 2000 Pezizales]
Asexual morph of Rhizostilbella type.

Ascocalathium Eidam ex J. Schröt.
in Cohn, Krypt.-Fl. Schlesien, 3.2 (1-2): 32 (1893)
Type species: Ascocalathium stipitatum Eidam ex J. Schröt. 1893
Family: Pyronemataceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Ascodesmis Tiegh.
Bull. Soc. bot. Fr., 23: 275 (1876)
Type species: Ascodesmis aurea Tiegh. 1876
Family: Ascodesmidaceae
Number of species: 6 fide DoF [7 integrated in 2000 Pezizales]

Ascophanus Boud.
Ann. Sci. Nat., Bot., sér. 5, 10: 241 (1869)
Type species: Ascophanus granuliformis (P. Crouan & H. Crouan) Boud. 1869
Synonym of Iodophanus ?

Ascorhizoctonia Chin S. Yang & Korf
Mycotaxon, 23: 468 (1985)
Type species: Ascorhizoctonia praecox Chin S. Yang & Korf 1985
Family: Pyronemataceae
Number of species: 7 fide DoF [1 integrated in 2000 Pezizales]

Ascosparassis Kobayasi
Bull. nat. Sci. Mus., Tokyo, n.s., 5: 45 (1960)
Type species: Ascosparassis shimizuensis Kobayasi 1960
Family: Otideaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Fructifications growing from underground sclerotioid carnose mycelial mass, stipitate coriaceous, flabellately dilated, margin lobed, undulate, entire; hymenium unilateral smooth; exciple (ectal excipulum) verrucose, parenchymatic; inner tissue (medullary excipulum) prosenchymatic. Paraphyses simple, not forming epithecium, apex peg-shaped; asci cylindric 8-spored inoperculate, apex opening by pore, not bluing with iodine. Ascospores uniseriate, ellipsoid smooth one-celled, hyaline.

Aurantiolachnea Van Vooren
Ascomycete.org, 13 (1): 9 (2021)
Type species: Aurantiolachnea solsequia (Quél.) Van Vooren 2021
Diagnosis: Differs from Humaria by the orange-coloured hymenium, smooth or finely dotted ascospores, its saprobic trophic status and genetic profile.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, orange-coloured, with an external surface and margin covered by brown hairs. Excipulum two-layered: medullary layer of textura intricata, ectal layer of textura subglobulosa/angularis. Hairs simple, superficial, septate, with a simple base. Asci operculate, narrowing towards the base, arising from croziers, inamyloid, 8-spored. Paraphyses slender, not enlarged at the top, containing carotenoid pigments. Ascospores uniseriate, ellipsoid, hyaline, biguttulate, smooth or very finely dotted. Trophic status probably saprobic. Asexual morph unknown.

Aurophora Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 52 (1968)
Type species: Aurophora dochmia (Berk. & M.A. Curtis) Rifai 1968
Family: Sarcoscyphaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Babosia D.G. Knapp, Zagyva, Trappe & Kovács
Mycologia, 112 (4): 811 (2020)
Type species: Babosia variospora D.G. Knapp, Zagyva, Trappe & Kovács 2020
Diagnosis: Ascomata gasteroid, hypogeous. Differs from the phylogenetically most closely related Temperantia, Hydnobolites, and Stouffera by the dark sporogenous zone and dark brown spore color at maturity, the clear peridermal layer at maturity, and the variability of the ornamentation of the ascospores.
Family: Pezizaceae
Number of species: 1 [0 integrated in 2000 Pezizales]

Balsamia Vittad.
Monogr. Tuberac.: 30 (1831)
Type species: Balsamia vulgaris Vittad. 1831
Family: Helvellaceae
Number of species: 6 fide DoF [5 integrated in 2000 Pezizales]

Barssia Gilkey
Mycologia, 17 (6): 253 (1925)
Type species: Barssia oregonensis Gilkey 1925
Family: Helvellaceae
Number of species: 2 fide DoF [6 integrated in 2000 Pezizales]

Berggrenia Cooke
Grevillea, 8: 64 (1879)
Type species: Berggrenia aurantiaca Cooke 1879
Family: Incertae sedis
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Boubovia Svrček
Česká Mykol., 31 (2): 71 (1977)
Type species: Boubovia luteola (Velen.) Svrček 1977
Family: Coprotaceae
Number of species: 6 fide DoF [2 integrated in 2000 Pezizales]

Boudiera Cooke
Grevillea, 6 (38): 76 (1877)
Type species: Boudiera areolata Cooke & W. Phillips 1877
Family: Pezizaceae
Number of species: 10 fide DoF [2 integrated in 2000 Pezizales]

Boudierella Sacc.
Bull. Acad. R. Sci. Belg. Cl. Sci., sér. 5, 34 (1): 130 (1895)
Type species: Boudierella coronata Costantin 1897
Family: Pyronemataceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Byssonectria P. Karst.
Meddn Soc. Fauna Flora fenn., 6: 6 (1881)
Type species: Byssonectria obducens P. Karst. 1881
Family: Pyronemataceae
Number of species: 10 fide DoF [4 integrated in 2000 Pezizales]

Caloscypha Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Caloscypha fulgens (Pers.) Boud. 1885
Family: Caloscyphaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Asexual morph of Geniculodendron type.

Carbomyces Gilkey
N. Amer. Fl. Ser., 2, 1: 27 (1954)
Type species: Carbomyces emergens Gilkey 1954
Family: Pezizaceae
Number of species: 3 fide DoF [1 integrated in 2000 Pezizales]

Cazia Trappe
Mem. N. Y. bot. Gdn, 49: 336 (1989)
Type species: Cazia flexiascus Trappe 1989
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Cephaliophora Thaxt.
Bot. Gaz., 35: 157 (1903)
Type species: Cephaliophora tropica Thaxt. 1903
Family: Ascodesmidaceae
Number of species: 6 fide DoF [0 integrated in 2000 Pezizales]

Chaetothiersia B.A. Perry & Pfister
Fungal Divers., 28: 69 (2008)
Type species: Chaetothiersia vernalis B.A. Perry & Pfister 2008
Family: Pyronemataceae
Number of species: 1 fide DoF [4 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, cupuliform, pale grey to whitish-coloured or yellow to orangish yellow, with an external surface covered by brown hairs. Margin hairy. Excipulum two-layered: medullary layer thick, of textura intricata, and ectal layer of textura globulosa/angularis. Hairs superficial, simple, septate, with a simple ± swollen base. Asci operculate, narrowing toward base, arising from free croziers, inamyloid, 8-spored. Paraphyses slender, equal to enlarged (sometimes subcapitate) at the top, hyaline or containing a sparse granular yellow pigment. Ascospores uniseriate, ellipsoid, hyaline, guttulate or not, smooth. Species saprobic on the litter and woody debris of conifers. Asexual morph unknown.

Chalazion Dissing & Sivertsen
Norw. J. Bot., 22 (1): 1 (1975)
Type species: Chalazion sociabile Dissing & Sivertsen 1975
Family: Ascodesmidaceae
Number of species: 3 fide DoF [3 integrated in 2000 Pezizales]

Cheilymenia Boud.
Bull. Soc. mycol. Fr., 1: 105 (1885)
Type species: Cheilymenia stercorea (Pers.) Boud. 1907
Family: Pyronemataceae
Number of species: 66 fide DoF [17 integrated in 2000 Pezizales]

Choiromyces Vittad.
Monogr. Tuberac.: 50 (1831)
Type species: Choiromyces meandriformis Vittad. 1831
Family: Tuberaceae
Number of species: 5 fide DoF [3 integrated in 2000 Pezizales]

Chorioactis Kupfer ex Eckblad
Nytt Mag. Bot., 15 (1-2): 127 (1968)
Type species: Chorioactis geaster (Peck) Kupfer ex Eckblad 1968
Family: Chorioactidaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Cidaris Fr.
Summa veg. Scand., sectio post.: 347 (1849)
Type species: Cidaris caroliniana (Schwein.) Fr. 1875
Family: Helvellaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Ciliaria Quél., nom. illeg.
Bull. Soc. bot. Fr., 26: 235 (1880)
Type species: Ciliaria fuscoatra (Rebent.) Quél. 1880
Synonym of Scutellinia

Cleistoiodophanus J.L. Bezerra & Kimbr.
Am. J. Bot., 63: 838 (1976)
Type species: Cleistoiodophanus conglutinatus J.L. Bezerra & Kimbr. 1976
Family: Ascobolaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Cleistothelebolus Malloch & Cain
Can. J. Bot., 49 (6): 851 (1971)
Type species: Cleistothelebolus nipigonensis Malloch & Cain 1971
Family: Pyronemataceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]
Current definition: Ascocarp initials coiled. Ascocarps subglobose to irregular in shape, pale brown, nonostiolate, nonstromatic. Asci irregularly disposed, ovoid, evanescent. Ascospores one-celled, hyaline, without germ pores, without "de Bary bubbles." Conidia produced as blastospores, hyaline, onecelled, often budding.

Complexipes C. Walker
Mycotaxon, 10 (1): 99  (1979)
Type species: Complexipes moniliformis C. Walker 1979
Family: Pyronemataceae
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Conoplea Pers.
Tent. disp. meth. fung.: 55  (1797)
Type species: Conoplea sphaerica (Pers.) Pers. 1797
Family: Sarcosomataceae
Number of species: 8 fide DoF [0 integrated in 2000 Pezizales]

Cookeina Kuntze
Revis. gen. pl., 2: 849  (1891)
Type species: Cookeina tricholoma (Mont.) Kuntze 1891
Family: Sarcoscyphaceae
Number of species: 12 fide DoF [8 integrated in 2000 Pezizales]

Coprobia Boud.
Bull. Soc. mycol. Fr., 1: 107 (1885)
Type species: Coprobia granulata (Bull.) Boud. 1904
Synonym of Cheilymenia

Coprotus Korf & Kimbr.
Am. J. Bot., 54 (1): 21 (1967)
Type species: Coprotus sexdecimsporus (P. Crouan & H. Crouan) Kimbr. & Korf 1967
Family: Coprotaceae
Number of species: 23 fide DoF [4 integrated in 2000 Pezizales]

Cubonia Sacc.
Syll. fung., 8: 527 (1889)
Type species: Cubonia brachyasca (Marchal) Sacc. 1889
Synonym of Ascobolus

Cupulina Dougoud, Van Vooren & M. Vega
Ascomycete.org, 7 (2): 39 (2015)
Type species: Cupulina montana Dougoud, Van Vooren & M. Vega 2015
Family: Pyronemataceae
Number of species: 2 [2 integrated in 2000 Pezizales]
Current definition: Apothecia gregarious, hemispherical or cupulate, sessile, partially embedded in the substrate or superficial, small, brownish ochraceous yellow, brownish orange or orange. Margin protruding, grossly and irregularly crenulate or fimbriate, whitish to pale yellow. Asci operculate, octospored, inamyloid. Ascospores fusoid or ellipsoid, hyaline. Paraphyses straight, enlarged at the top, with yellowish or hyaline content, without iodine reaction, septate, simple or furcate. Medullary excipulum of textura intricata. Ectal excipulum of textura globulosa to globulosa-angularis. Habitat: on soil, on naked or mossy ground.

Cyathipodia Boud.
Hist. Class. Discom. Eur.: 39 (1907)
Type species: Cyathipodia platypodia Boud. 1907
Synonym of Helvella

Daleomyces Setch.
Mycologia, 16 (5): 240 (1924)
Type species: Daleomyces gardneri Setch. 1924
Family: Pezizaceae
Number of species: 6 [4 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, cupulate, discoid or sparassoid (cabbage-head), of various colours: whitish with pinkish tinges, becoming more or less brownish with age, purplish brown, olivaceous brown or distinctly violet. Flesh without latex. Asci operculate, 8-spored, with the top strongly bluing in an iodine solution (WTR type), with or without crozier. Paraphyses containing a vacuolar pigment, brown, purple brown or yellowish becoming brown with maturity. Ectal excipulum of textura globulosa, with large cells, sometimes partly gelatinised in the outer part (e.g. D. exogelatinosus). Ascospores biguttulate, ornamented with isolate warts, punctiform or irregular, often more dense at the poles. Species saprobic, preferentially growing on basic substrates (burned soil, sandy soil, soil with a high pH).

Delastria Tul. & C. Tul.
Ann. Sci. Nat. Bot., sér. 2, 19: 379  (1843)
Type species: Delastria rosea Tul. & C. Tul. 1843
Family: Pezizaceae
Number of species: 1 fide DoF [6 integrated in 2000 Pezizales]

Dennisiopsis Subram. & Chandrash.
Kew Bull., 31 (3): 639 (1977)
Type species: Dennisiopsis octospora Subram. & Chandrash. 1977
Family: Incertae sedis
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Densocarpa Gilkey
N. Amer. Fl., ser. 2, 1: 15 (1954)
Type species: Densocarpa shanorii Gilkey 1954
Family: Tarzettaceae
Number of species: 2 [2 integrated in 2000 Pezizales]

Desmazierella Lib.
Ann. Sci. Nat. Bot., sér. 1, 17: 83  (1829)
Type species: Desmazierella acicola Lib. 1829
Family: Chorioactidaceae
Number of species: 2 fide DoF [2 integrated in 2000 Pezizales]
Asexual morph of Verticicladium type.

Dichobotrys Hennebert
Persoonia, 7 (2): 193 (1973)
Type species: Dichobotrys abundans Hennebert 1973
Synonym of Trichophaea

Dictyocoprotus J.C. Krug & R.S. Khan
Mycologia, 83 (1): 103  (1991)
Type species: Dictyocoprotus mexicanus J.C. Krug & R.S. Khan 1991
Family: Ascodesmidaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Apothecla scattered, sessile, smooth, discoid to saucer-shaped, whitish to pale yellowish; excipulum cyanophilous, non-dextrinoid, two-layered, composed of an ectal excipulum of textura angularis and a distinct medullary excipulum of textura intricata. Asci protruding, operculate, nonamyloid, cylindrical, tapering below into a short stipe. Paraphyses filiform, simple, hyaline, enlarged and uncinate at the apices. Ascospores one-celled, ellipsoidal, hyaline to pale yellowish, ornamented with a fine reticulum, at maturity forming de Bary bubbles, lacking mucilaginous sheath.

Diehliomyces Gilkey
Mycologia, 46 (5): 789 (1954)
Type species: Diehliomyces microsporus (Diehl & E.B. Lamb.) Gilkey 1954
Family: Otideaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Dingleya Trappe
Mycotaxon, 9 (1): 331  (1979)
Type species: Dingleya verrucosa Trappe 1979
Family: Tuberaceae
Number of species: 7 fide DoF [3 integrated in 2000 Pezizales]
Current definition: Ascocarps hypogeous with a brown, verrucose peridium. Gleba white with rounded to narrow, labyrinthine chambers lined with a hymenial palisade and filled with asci, paraphyses and spores. Asci cylindric, mostly 8-spored. Spores globose to ellipsoid, hyaline to brown, ornamented with cyanophilic, rounded to irregular warts or ridges.

Discina (Fr.) Fr.
Summa veg. Scand., section post.: 348  (1849)
Type species: Discina perlata (Fr.) Fr. 1849
Synonym of Gyromitra

Discinella P. Karst., nom. illeg.
Hedwigia, 30: 301 (1891)
Type species: Discinella corticalis P. Karst. 1891
Family: Incertae sedis
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Disciotis Boud.
Bull. Soc. mycol. Fr., 1: 100  (1885)
Type species: Peziza venosa Pers. 1801
Family: Morchellaceae
Number of species: 4 fide DoF [1 integrated in 2000 Pezizales]

Dissingia K. Hansen, X.H. Wang & T. Schumach.
Persoonia, 42: 195 (2019)
Type species: Dissingia leucomelaena (Pers.) K. Hansen & X.H. Wang 2019
Family: Helvellaceae
Number of species: 4 [4 integrated in 2000 Pezizales]
Current definition: Apothecia cup-shaped, mostly remaining concave when expanding, subsessile, or with a ± distinct stipe; stipe short, broad or slender, with a few ± conspicuous, blunt ribs gradually widening and subdividing towards the cup attachment. Hymenium yellowish brown to greyish brown, to dark brownish black; receptacle surface even or with few to many blunt to angular ribs and grooves below, pubescent, upper part whitish to dark greyish brown, below concolorous or often gradually paler to almost whitish. Asci cylindrical, operculate, arising from simple septa, 8-spored. Spores ellipsoid, obtuse or attenuate, smooth. Paraphyses filiform, septate, with clavate or subcapitate tips.

Donadinia Bellem. & Mel.-Howell
Cryptog. Mycol., 11 (3): 218  (1990)
Type species: Donadinia helvelloides (Donadini, Berthet & Astier) Bellem. & Mel.-Howell 1990
Diagnosis: Apotheciae lignicolae, in ligno mortuo caduco enascentes, pedunculatae, cupulares, gregariae; cupula hemisphaerica, marginata. Stipes plenus. Asci octospori, cylindrati, base flexuosi. Endotunica in apice solum adest, natura heteroclita. Ectotunica in apice angustata, natura heteroclita. Apertio operculata haud suboperculata. Sporae pariete propra dispari, ae perispora verrucis parvis cyanophilisque ornata, et intus parcis guttulis repletae. Ascogenae hyphae non fibulatae. Paraphyses lineares, apice saepe furcatae ac nodosae.
Family: Sarcosomataceae
Number of species: 1 fide DoF [5 integrated in 2000 Pezizales]

Durandiomyces Seaver
N. Amer. Cup-Fungi (Operc.): 242 (1928)
Type species: Durandiomyces phillipsii (Massee) Seaver 1928
Synonym of Daleomyces

Elaiopezia Van Vooren
Ascomycete.org, 12 (4): 185 (2020)
Type species: Elaiopezia polaripapulata (J. Moravec) Van Vooren 2020
Family: Pezizaceae
Number of species: 5 [2 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, discoid or slightly cupulate, small-sized, olive or yellowish with distinct olivaceous tinges. Flesh without latex. Asci operculate, 8-spored; ascus wall diffusely amyloid (W type). Paraphyses containing a yellow, yellow-ochre or olivaceous pigment, sometimes also with an external pigment at the top. Ectal excipulum of textura globulosa with small to medium-sized cells. Ascospores eguttulate, sometimes containing small granules or drops in young state, smooth or ornamented. Species saprobic.

Eleutherascus Arx
Persoonia, 6 (3): 377  (1971)
Type species: Eleutherascus lectardii (Nicot) Arx 1971
Family: Ascodesmidaceae
Number of species: 4 fide DoF [1 integrated in 2000 Pezizales]

Eoaleurina Korf & W.Y. Zhuang
Mycotaxon, 26: 381  (1986)
Type species: Eoaleurina foliicola (Henn.) Korf & W.Y. Zhuang 1986
Family: Pyronemataceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Eoterfezia G.F. Atk.
Bot. Gaz., 34: 40 (1902)
Type species: Eoterfezia parasitica G.F. Atk. 1902
Family: Incertae sedis
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Eremiomyces Trappe & Kagan-Zur
Mycol. Res., 109 (2): 244 (2005)
Type species: Eremiomyces echinulatus (Trappe & Marasas) Trappe & Kagan-Zur 2005
Diagnosis: A Choiromyces sporis virgis conicis rectis obtusis, cellulis magnopere inflatis, ordiniibus DNA abhorrentibus (ad Pezizaceas, non Tuberaceas, affinis) et distributione in desertis Hemisphaerio Meridionali differt.
Family: Pezizaceae
Number of species: 1 fide DoF [3 integrated in 2000 Pezizales]

Fallacidiscus U. Lindemann, Roffler & Van Vooren
Ascomycete.org, 14 (2): 46 (2022)
Type species: Fallacidiscus helveticus U. Lindemann, Roffler & Van Vooren 2022
Diagnosis: Differs from Spaniodiscus fuckelii and Ramsbottomia spp. by discoid apothecia with a small marginal edge, a hairless margin, and finely warted ascospores containing polar oil droplets, as well as its genetic profile.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, yellow-orange-coloured, with an external surface densely covered by short brown hairs and a hairless margin. Excipulum two-layered: medullary layer of textura intricata, ectal layer of textura subglobulosa/angularis. Hairs short, simple, superficial, septate, with a simple base, obtuse or slightly sharp. Asci operculate, 8-spored, narrowing toward base, arising from croziers, inamyloid. Paraphyses slender, ± enlarged at the top, containing carotenoid pigments. Ascospores uniseriate, ellipsoid, hyaline, thick-walled, containing small bipolar oil droplets, finely warted. Trophic status unknown. Asexual morph unknown.

Fastigiella Benedix
Kulturpflanze, 17: 276 (1969)
Type species: Fastigiella caroliniana (Bosc) Benedix 1969
Synonym of Gyromitra

Filicupula Y.J. Yao & Spooner
Kew Bull., 51 (1): 193  (1996)
Type species: Filicupula suboperculata (Döbbeler & P. James) Y.J. Yao & Spooner 1996
Family: Incertae sedis
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Apothecia clavate at first, becoming obovoid or turbinate, flattened above. Disc orange. Receptacle cupulate, surface rough, with branched anchoring hyphae present at the base and lower receptacle. Ectal excipulum comprising textura prismatica at the base, and septate, sub-parallel hyphae above; excipular hyphae radially arranged, lying at a low angle to the surface, their walls thickened and strongly adherent. Asci operculate, iodine-negative, clavate, thin-walled. Ascospores unicellular, colourless, smooth, fusoid, often inaequilateral, guttulate. Paraphyses filiform, septate, straight or curved, flexuous.

Fischerula Mattir.
Nuovo G. bot. ital., 34: 1348  (1928)
Type species: Fischerula macrospora Mattir. 1928
Family: Morchellaceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]

Fuckelina Kuntze
Rev. Gener. Plant., 3: 852 (1891)
Type species: Fuckelina macropus (Pers.) Kuntze 1891
Synonym of Helvella

Galactinia (Cooke) Boud.
Bull. Soc. mycol. Fr., 1: 101  (1885)
Type species: Galactinia saniosa (Schrad.) Sacc. 1889
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Galeoscypha Svrček & J. Moravec
Česká Mykol., 43 (4): 210  (1989)
Type species: Galeoscypha pileiformis (Svrček) Svrček & J. Moravec 1989
Diagnosis: Carposomata parva, gregaria, usque confluentia, superficialia, absque hypothallo sed basi hyphis ferrugineo-brunneis, tenuibus, crasse tunicatis, septatis, dense connexis substrato adnata, primum subglobosa, pyriformia, apice truncata, dein pileiformia, disco convexo, hemisphaerico, immarginato, nonnumquam obtuse conico, margine undulato, granuloso, fulgide aurantiaco, subtus crasse obconica vel subcylindrica, stipitiformiter angustata, thecio sanguineo-rubro vel aurantiaco-rubro, extus minutissime granulata Excipulum externum atque medullaris textura globuloso-angularis, aurantiaco-coloratum, parte externa margineque cellulis globosis cyanophilis, ad basim pilis hyphoideis flexuosis hyalinis septatis et hyphis mycelialibus ferrugineo-brunneis instructum. Asci cylindrici, operculati, non amyloidei, octospori. Paraphyses crasse cylindricae, parte basali ramosae, sursum dilatatae, clavatae, subcapitatae vel lanceolatae, septatae, sub hyalinae. Ascosporae ellipsoideae, inaequales, intus irregulariter granulosae sed etiam biguttulatae, perisporio non separabile, verrucis cyanophilis in aequalibus ornato. Habitat saprobioticus.
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Galiella Nannf. & Korf
Mycologia, 49 (1): 107  (1957)
Type species: Galiella rufa (Schwein.) Nannf. & Korf 1957
Family: Sarcosomataceae
Number of species: 8 fide DoF [2 integrated in 2000 Pezizales]

Genabea Tul. & C. Tul.
G. bot. ital., 1 (7-8): 60  (1845)
Type species: Genabea fragilis Tul. & C. Tul. 1845
Family: Pyronemataceae
Number of species: 1 fide DoF [3 integrated in 2000 Pezizales]

Genea Vittad.
Monogr. Tuberac.: 27  (1831)
Type species: Genea verrucosa Vittad. 1831
Family: Pyronemataceae
Number of species: 32 fide DoF [17 integrated in 2000 Pezizales]

Geneosperma Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 102 (1968)
Type species: Geneosperma geneosporum Rifai 1968
Family: Pyronemataceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Geodina Denison
Mycologia, 57 (4): 649  (1965)
Type species: Geodina guanacastensis Denison 1965
Family: Wynneaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Apothecium goblet-shaped with a slender stem, thick-fleshed, corky to leathery; hymenium light-colored to bright-colored, yellow to red; exterior similarly colored, hairy; hairs fasciculate composed of thick-walled, nonseptate hyphae arising from the ectal excipulum; excipulum composed of two distinct layers, a medullary excipulum of entangled hyphae (textura intricata), and a rind-like ectal excipulum composed of more or less rectangular cells (textura prismatica to textura angularis); asci suboperculate, with eccentric opercula, cylindrical with gradually tapering bases, not turning blue in iodine, not all maturiilg simultaneously; ascospores ellipsoid, large, long, containing 1-2 large oil drops, heavily sculptured; spore sculpturing not staining readily in cotton blue, consisting of coarse, anastomosing ridges; paraphyses slender, cylindrical, septate, with subclavate apices; growing in the soil in the tropics.

Geomorium Speg.
Anal. Soc. cient. argent., 94 (1-2): 79 (1922)
Type species: Geomorium fuegianum Speg. 1922
Family: Geomoriaceae
Number of species: 8 [6 integrated in 2000 Pezizales]

Geopora Harkn.
Bull. Calif. Acad. Sci., 1 (3): 168  (1885)
Type species: Geopora cooperi Harkn. 1885
Family: Pyronemataceae
Number of species: 13 fide DoF [7 integrated in 2000 Pezizales]

Geopyxis (Pers.) Sacc.
Syll. fung. (Abellini), 8: 63  (1889)
Type species: Geopyxis carbonaria (Alb. & Schwein.) Sacc. 1889
Family: Tarzettaceae
Number of species: 7 fide DoF [6 integrated in 2000 Pezizales]

Geoscypha (Cooke) Lambotte
Mém. Soc. roy. sci. Liège, 2e sér., 14: 320 (1888)
Type species: Geoscypha violacea (Pers.) Lambotte 1888
Family: Pezizaceae
Number of species: 3 [4 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, cupulate or discoid, violet-coloured. Asci operculate, 8-spored, with the top intensely bluing in iodine solution (WTR or WT type in G. ampelina), with crozier. Paraphyses containing a brown or purple brown vacuolar pigment. Ectal excipulum of textura globulosa, with large cells. Ascospores smooth or ornamented with isolated warts, biguttulate, except in G. violacea where the drops may be visible only in young spores. Species saprobic, often growing on burnt soil.

Gilkeya M.E. Sm., Trappe & Rizzo
Mycologia, 98 (5): 705 (2007)
Type species: Gilkeya compacta (Harkn.) M.E. Sm. & Trappe 2007
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Glaziella Berk.
Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn, 80: 31  (1880)
Type species: Glaziella vesiculosa Berk. 1880
Family: Glaziellaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Globopilea Beauseign.
Contr. Etude Fl. Mycol. Landes: 204 (1926)
Type species: Globopilea phlebophora (Pat. & Doass.) Beauseign. 1926
Synonym of Helvella

Greletia Donadini
Bull. Soc. mycol. Fr., 95(3): 183 (1980)
Type species: Greletia planchonis (Dunal ex Boud.) Donadini 1980
Synonym of Smardaea

Gymnohydnotrya B.C. Zhang & Minter
Mycol. Res., 92 (2): 192  (1989)
Type species: Gymnohydnotrya australiana B.C. Zhang & Minter 1989
Synonym of Geomorium

Gyrocephalus Pers., nom. rejic.
Mém. Soc. Linn. Paris, 3: 77 (1824)
Type species: Gyrocephalus aginnensis Pers. 1824
Synonym of Gyromitra

Gyromitra Fr.
Summa veg. Scand., section post.: 346  (1849)
Type species: Gyromitra esculenta (Pers.) Fr. 1849
Family: Discinaceae
Number of species: 18 fide DoF [21 integrated in 2000 Pezizales]

Gyromitrodes Vassilkov
Sovetsk. Bot., 6: 51 (1942)
Type species: Gyromitrodes sphaerospora (Peck) Vassilkov 1942
Synonym of Gyromitra

Hansenopezia Matočec, I. Kušan & Jadan
in Yuan et al., Fungal Divers., 104 : 64 (2020)
Type species: Hansenopezia retrocurvata (K. Hansen & Sandal) Matočec, I. Kušan & Jadan 2020
Family: Pezizaceae
Number of species: 2 [2 integrated in 2000 Pezizales]
Current definition: Ascomata apothecial, solitary or often in groups, medium-sized, epigeous, centrally attached to the substrate, regularly circular to much sinuous from the top view, cupulate when young, then plate-shaped, hymenial surface reddish brown, lilac-violet, violet-brown, purplish brown to dark brown, matte, margin upright then flattened to reflexed, entire, blunt, smooth and or beset with scattered white fibrils, whitish grey or pale brownish yellow, excipular ground concolourous with the margin, finely pubescent or ornamented with reddish brown patches distributed to the basal parts. Flesh fragile, homogenous, soft waxy, dry on cut or exuding watery to milk-white latex. Hymenium arranged as a regular palisade. Asci cylindrical, hyaline, apex subtruncate, protruding above paraphysis tips at full maturity, 8-spored, base tapered, arising from croziers, operculum apical, very narrow, occupying ≤ 28% of living mature ascus diam, functional, flat lentiform before opening, encircled by slight “u”-shaped indentation in the wall, periascal amyloid mucus thin and film-like through the entire length but almost lacking on opercular lense, while the thickest at the opercular edge where is moderately euamyloid, forming unclear thin annular zone, reaction gradually decreasing in strength towards the base, with reacting area on half of the ascus length or more. Ascospores hyaline, ellipsoid to narrowly ellipsoid with ± rounded poles, radially symmetrical, 1-celled, subglabrous or with cyanophilic verrucae, perispore cyanophilic, not loosening when heated in lactic Cotton Blue or in KOH, wall 3-layered ± thick-walled, without sheath on ejected spores; sporoplasm multiguttulate, lipid content nearly maximal with all guttules of approx. same size or 2–3 large lipid bodies mixed with many smaller ones, lipid bodies much coalesced in dead spores, uninucleate, nucleus and nucleolus not contrasted nor stained in Lugol’s solution, de Bary bubbles absent in anhydrous media. Paraphyses cylindrical, apically obtuse to subclavate, straight to bent, not branching in the upper part, embedded in hyaline epithecial exudate or only with hyaline, thin coating, apical cell contain single or few non-refractive vacuoles entirely filled with finely dispersed minute pinkish brown particles and/or partially with larger non-refractive granules of the same colour, not containing vacuolar bodies, in Lugol’s solution pigment partially turning to rusty brown colour. Marginal texture in section displaying broad outline, textura globulosa-angularis mixed with some cylindric cell chains that may protrude as scattered cylindric obtuse, thin-walled hyphoid hairs or form loose tufts; texture is not cyanophilous. Subhymenium well-discerned, composed of hyaline subgelatinous densely woven textura porrecta. Medullary and ectal excipulum of ± homogenous, subhyaline texture, except for the cortical layer. Upper part composed of textura globulosa- moniliformis mixed with cylindric very long, mostly vertically oriented hyphae. Lower part composed of textura globulosa-angularis with intermixed intricate connective hyphae, cells smaller in size towards the surface. Cortical layer composed of textura porrecta-intricata, discontinuous layer when present pustulate, cells with ochraceous walls and rusty brown patches, coiled and agglutinated together; texture is not cyanophilous.
Ecology: Saprotroph on rich soil with litter and wood residues of Abies alba, on very rotten stumps and trunks of Picea abies and Fagus sylvatica, sometimes also on rich soil outside forests, fruitbodies appear in spring or autumn to winter.

Hapsidomyces J.C. Krug & Jeng
Mycologia, 76 (4): 748  (1984)
Type species: Hapsidomyces venezuelensis J.C. Krug & Jeng 1984
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Hellenicoscyphus U. Lindemann, Van Vooren & Kaounas
Ascomycete.org, 14 (2): 74 (2022)
Type species: Hellenicoscyphus hyalotrichus U. Lindemann, Van Vooren & Kaounas 2022
Diagnosis: Differs from Paratricharina by its hyaline marginal hairs organised in small bundles, the lack of excipular hairs and from Tricharina by its hyaline marginal hairs, the lack of excipular hairs and its bipolar spore granules, and from both genera by its genetic profile.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, cupuliform, light brown to ochre. Receptacle hairless. Marginal hairs organised in small bundles, superficial, septate, with a simple widened base. Excipulum two-layered: medullary layer of textura intricata, and ectal layer of textura globulosa/angularis. Asci operculate, narrowing toward base, arising from free croziers, inamyloid, 8-spored. Paraphyses slender, hyaline. Ascospores uniseriate, ellipsoid, hyaline, smooth, containing small bipolar granules. Species probably saprobic. Asexual morph unknown.

Helvella L.
Sp. pl., 2: 1180  (1753)
Type species: Helvella mitra L. 1753
Family: Helvellaceae
Number of species: 52 fide DoF [56 integrated in 2000 Pezizales]

Helvellella S. Imai
Bot. Mag. (Tokyo), 46: 174 (1932)
Type species: Helvellella sphaerospora (Peck) S. Imai 1932
Synonym of Gyromitra

Heteroplegma Clem.
Bull. Torrey bot. Club, 30: 92 (1903)
Type species: Heteroplegma caeruleum Clem. 1903
Synonym of Peziza

Heydenia Fresen.
Beitr. Mykol., 2: 47 (1852)
Type species: Heydenia alpina Fresen. 1852
Family: Pseudombrophilaceae
Number of species: 4 fide DoF [2 integrated in 2000 Pezizales]

Hiemsia Svrček
Česká Mykol., 23: 83  (1969)
Type species: Hiemsia pseudoampezzana (Svrček) Svrček 1969
Family: Pyronemataceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Hoffmannoscypha Stielow, Göker & Klenk
Mycol. Progr., 12 (4): 684 (2012)
Type species: Hoffmannoscypha pellita (Sacc.) Stielow, Göker & Klenk 2012
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Apothecia superficially attached to the substrate, often slightly sunken into it, without stipe, roundish when juvenile and nearly entirely closed at the apex, cupulate when mature, at full maturity with ripped edges; lobes erect when juvenile, effused when mature. Ectal excipulum (epicutis) dark yellow-orange, seldom orange-brown, pseudoparenchymatous, with angular or isodiametric, thick-walled cell agglomerates (textura angularis), cells giving rise to dark brown, thick-walled, multiple septate cylindrical finely warted setae, single or cespitose, dark brown in water and 5% KOH. Asci operculate, cylindrical, eight-spored, monoseriate. Paraphyses slender, single, septate, with gentle orange pigmentation giving rise to the colour of the hymenium, slightly thickened and bent at the tip, arranged in palisade order.

Humaria Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 320  (1870)
Type species: Humaria leucoloma (Hedw.) Sacc. 1889
Family: Pyronemataceae
Number of species: 16 fide DoF [1 integrated in 2000 Pezizales]

Hydnobolites Tul. & C. Tul.
Ann. Sci. Nat. Bot., sér. 2, 19: 278  (1843)
Type species: Hydnobolites cerebriformis Tul. & C. Tul. 1843
Family: Pezizaceae
Number of species: 3 fide DoF [4 integrated in 2000 Pezizales]

Hydnocystis Tul. & C. Tul.
G. bot. ital., 1 (7-8): 59  (1845)
Type species: Hydnocystis arenaria Tul. & C. Tul. 1845
Family: Tarzettaceae
Number of species: 2 fide DoF [4 integrated in 2000 Pezizales]

Hydnoplicata Gilkey
Mycologia, 46 (5): 783 (1955)
Type species: Hydnoplicata whitei Gilkey 1954
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Hydnotrya Berk. & Broome
Ann. Mag. nat. Hist., ser. 1, 18: 78  (1846)
Type species: Hydnotrya tulasnei (Berk.) Berk. & Broome 1846
Family: Discinaceae
Number of species: 15 fide DoF [9 integrated in 2000 Pezizales]

Hydnotryopsis Gilkey
University of Calif. Publ. Bot., 6: 336  (1916)
Type species: Hydnotryopsis setchellii Gilkey 1916
Family: Pezizaceae
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Hypotarzetta Donadini
Docums Mycol., 15 (60): 48:  (1985)
Type species: Hypotarzetta insignis (Berthet & Riousset) Donadini 1985
Synonym of Tarzetta

Imaia Trappe & Kovács
Mycologia, 100 (6): 934 (2008)
Type species: Imaia gigantea (S. Imai) Trappe & Kovács 2008
Family: Morchellaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Infundibulum Velen.
Monogr. Discom. Bohem.: 351 (1934)
Type species: Infundibulum linteicola (W. Phillips & Plowr.) Velen. 1934
Synonym of Peziza

Iodophanus Korf
Am. J. Bot., 54 (1): 18  (1967)
Type species: Iodophanus carneus (Pers.) Korf 1967
Family: Pezizaceae
Number of species: 16 fide DoF [1 integrated in 2000 Pezizales]
Asexual morph of Oedocephalum type.

Iodowynnea Medel, Guzmán & S. Chacón
Mycotaxon, 59: 128  (1996)
Type species: Iodowynnea auriformis (Pat. ex Le Gal) Medel, Guzmán & S. Chacón 1996
Diagnosis: A Wynnea affine differt ascis iodinos et sporae verrucosis in vittas in longitudinem porrectas confluentibus.
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Ionopezia Van Vooren
Ascomycete.org, 12 (4): 187 (2020)
Type species: Ionopezia gerardii (Cooke) Van Vooren 2020
Family: Pezizaceae
Number of species: 2 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, discoid or cupulate, sessile, violet-coloured. Flesh without latex. Asci operculate, 8-spored, with or without crozier, wall diffusely amyloid (W type). Paraphyses hyaline, with an external purplish brown to violet pigment located at the top. Ascospores fusoid, multiguttulate, ornamented with longitudinal striations (only seen with a high magnification). Species forming ectomycorrhizas.

Iotidea Clem.
Gen. fung.: 89 (1909)
Type species: Iotidea pleurota (W. Phillips) Clem. 1909
Synonym of Peziza ?

Jafnea Korf
Nagaoa, 7: 5  (1960)
Type species: Jafnea fusicarpa (W.R. Gerard) Korf 1960
Family: Pyronemataceae
Number of species: 2 fide DoF [3 integrated in 2000 Pezizales]

Kalaharituber Trappe & Kagan-Zur
Mycol. Res., 109 (2): 242 (2005)
Type species: Kalaharituber pfeilii (Henn.) Trappe & Kagan-Zur 2005
Diagnosis: A Terfezia sporis spinis minutissimis dense congestis, ordiniibus DNA abhorrentibus, distributione in Hemisphaerio Meridionali differt.
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Kalapuya M.J. Trappe, Trappe & Bonito
Mycologia, 102 (5): 1059 (2010)
Type species: Kalapuya brunnea M.J. Trappe, Trappe & Bonito 2010
Diagnosis: Ascomata hypogaea, stereothecia, subglobosa. Peridium porphyreum vel brunneum, verrucosum, verrucis fissuris tenuibus separatis. Gleba solida, albida, contextu fertili griseo-brunneo maculato. Asci ellipsoidei vel globosi. Sporae ellipsoideae, laeves, juventute hyalinae, maturitate succineae.
Family: Morchellaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Kallistoskypha Pfister, Agnello, Lantieri & LoBuglio
Mycol. Progr., 12 (4): 670 (2013)
Type species: Kallistoskypha incarnata (Duvernoy & Maire) Pfister, Agnello, Lantieri & LoBuglio 2013
Family: Caloscyphaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Karstenella Harmaja
Karstenia, 9: 20  (1969)
Type species: Karstenella vernalis Harmaja 1969
Family: Karstenellaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Ascocarp very thin, totally tightly appressed at the substrate (not being a true apothecium), its development probably gymnocarpic. Spores one-celled, rather thin-walled, hyaline in KOH, slightly yellowish in Melzer''s, blue in Cotton Blue (Methyl Blue) in lactic acid. Asci with persistent wall of the unitunicate type, discharging their spores with a puff, operculate, rather thin-walled, apex not differentiated, cylindrical, eight-spored, not blueing with Melzer''s. Excipulum throughout composed of more or less interwoven, filamentous hyphae (textura intricata). Subiculum (?) composed of interwoven, filamentous hyphae of other kind than those of the excipulum.

Kimbropezia Korf & W.Y. Zhuang
Mycotaxon, 40: 272 (1991)
Type species: Kimbropezia campestris Korf & W.Y. Zhuang 1991
Family: Pezizaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Kompsoscypha Pfister
Mem. N. Y. bot. Gard., 49: 340  (1989)
Type species: Kompsoscypha chudei (Pat. ex Le Gal) Pfister 1989
Family: Sarcoscyphaceae
Number of species: 4 fide DoF [3 integrated in 2000 Pezizales]

Korfiella D.C. Pant & V.P. Tewari
Trans. Br. mycol. Soc., 54 (3): 493  (1970)
Type species: Korfiella karnika D.C. Pant & V.P. Tewari 1970
Family: Sarcosomataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Kotlabaea Svrček
Česká Mykol., 23: 85  (1969)
Type species: Kotlabaea deformis (P. Karst.) Svrček 1969
Synonym of Byssonectria

Labyrinthomyces Boedijn
Bull. Jard. bot. Buitenz, sér. 3, 16 (2): 238  (1939)
Type species: Labyrinthomyces steenisii Boedijn 1939
Family: Tuberaceae
Number of species: 7 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Ascocarps hypogeous with a brown tomentose peridium. Gleba with prominent, labyrinthine, mostly hollow chambers that do not open to the surface. Asci cylindric, long-stipitate, mostly 8-spored. Paraphyses much shorter than asci. Spores globose, hyaline, smooth in youth, at maturity ornamented with rounded, irregular warts.

Lamprospora De Not.
Comm. Soc. crittog. Ital., 1 (5): 388  (1863)
Type species: Lamprospora miniata (P. Crouan & H. Crouan) De Not. 1864
Family: Pyronemataceae
Number of species: 46 fide DoF [28 integrated in 2000 Pezizales]

Lasiobolidium Malloch & Cain
Can. J. Bot., 49 (6): 853  (1971)
Type species: Lasiobolidium spirale Malloch & Cain 1971
Family: Pyronemataceae
Number of species: 7 fide DoF [6 integrated in 2000 Pezizales]
Current definition: Ascomata cleistothecial or apothecial, small, sessile, subglobose, slightly cupuliform or discoid, whitish, pale greyish or yellow-brown, with an external surface covered by brown hairs. Margin hairy on the apothecial species. Ectal excipulum of textura globulosa/angularis. Hairs ± superficial or deeply rooted, septate, with a simple base, sometimes multifurcate, long, upward-pointing, straight or helically coiled. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 4- or 8-spored. Ascospores ellipsoid, hyaline, eguttulate, but sometimes containing small granules usually disappearing in mature ascospores, smooth or verrucose with isolated warts. Species saprobic, coprophilous or not. Asexual morph unknown.

Lasiobolus Sacc.
Bot. Zbl., 18: 220  (1884)
Type species: Lasiobolus pilosus (Fr.) Sacc. 1884
Family: Ascodesmidaceae
Number of species: 11 fide DoF [5 integrated in 2000 Pezizales]

Lasiocupulina Van Vooren & M. Vega
Ascomycete.org, 10 (6): 221 (2018)
Type species: Lasiocupulina mediterranea Van Vooren & M. Vega 2018
Diagnosis: Differs from Cupulina by the presence of a hairy margin and its molecular data.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Lathraeodiscus Dissing & Sivertsen
Mycologia, 80 (6): 832  (1989)
Type species: Lathraeodiscus arcticus Dissing & Sivertsen 1989
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Lazuardia Rifai
Mycotaxon, 31 (1): 241  (1988)
Type species: Lazuardia lobata (Berk. & M.A. Curtis) Rifai 1988
Family: Pulvinulaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Legaliana Van Vooren
Ascomycete.org, 12 (4): 188 (2020)
Type species: Legaliana badia (Pers.) Van Vooren 2020
Family: Pezizaceae
Number of species: 6 [4 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, cupulate or discoid, sessile, dark coloured (brown, reddish brown, olivaceous brown, purplish brown, blackish purple). Flesh without coloured latex, but sometimes with a watery juice. Asci operculate, 8-spored, with crozier, wall diffusely bluing in an iodine solution, more or less intensely at the top (WT type). Paraphyses containing a brownish or olive-brown vacuolar pigment, and often showing an external brown pigment at the top. Ascospores biguttulate, sometimes with an oil drop larger than the other, more rarely uniguttulate, ornamented with irregular warts, but often with elongated warts or crests, sometimes forming a network or an incomplete reticulum. Ectal excipulum of textura globulosa/angularis, with large cells. Species saprobic or ectomycorrhizal.

Lepidotia Boud.
Bull. Soc. mycol. Fr., 1: 101 (1885)
Type species: Lepidotia hispida (Quél.) Boud. 1907
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Leptopodia Boud.
Bull. Soc. mycol. Fr., 1: 99 (1885)
Type species: Leptopodia elastica (Bull.) Boud. 1904
Synonym of Helvella

Leucangium Quél.
Compt. Rend. Assoc. Franç. Avancem. Sci., 11: 404  (1883)
Type species: Leucangium ophthalmosporum Quél. 1883
Family: Morchellaceae
Number of species: 1 fide DoF [2 integrated in 2000 Pezizales]

Leucopezis Clem.
Gen. Fungi: 175 (1909)
Type species: Leucopezis excipulata Clem. 1909
Synonym of Humaria ?

Leucoscypha Boud.
Bull. Soc. mycol. Fr., 1: 104  (1885)
Type species: Leucoscypha leucotricha (Alb. & Schwein.) Boud. 1885
Family: Pyronemataceae
Number of species: 10 fide DoF [1 integrated in 2000 Pezizales]

Loculotuber Trappe, Parladé & I.F. Alvarez
Mycologia, 84 (6): 927  (1993)
Type species: Loculotuber gennadii (Chatin) Trappe, Parladé & I.F. Alvarez 1993
Family: Incertae sedis
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Lotinia Pérez-Butrón, Fern.-Vic. & P. Alvarado
in Crous et al., Persoonia, 35: 325 (2015)
Type species: Lotinia verna Pérez-Butrón, Fernández-Vicente & P. Alvarado 2015
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Luciotrichus R. Galán & Raitv.
Czech Mycol., 47 (4): 272  (1995)
Type species: Luciotrichus lasioboloides R. Galán & Raitv. 1995
Family: Ascodesmidaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Luteoamylascus Cabero, P. Alvarado & G. Moreno
Mycol. Progr., 15 (4): 3 (2016)
Type species: Luteoamylascus aculeatus Cabero, P. Alvarado & G. Moreno 2016
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Macropodia Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 331 (1870)
Type species: Macropodia macropus (Pers.) Fuckel 1870
Synonym of Helvella

Macroscyphus Nees ex Gray
Nat. Arr. Brit. Plant., 1: 671 (1821)
Type species: Macroscyphus macropus (Pers.) Gray 1821
Synonym of Helvella

Malvipezia Van Vooren
Ascomycete.org, 12 (4): 188 (2020)
Type species: Malvipezia howsei (Roze & Boud) Van Vooren 2020
Family: Pezizaceae
Number of species: 4 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, cupulate or discoid, sessile, with pale mauve, purple or violet colours. Flesh without latex. Asci operculate, 8-spored, with crozier, wall diffusely amyloid in an iodine solution except at the top where the reaction is intense and forms a ring (WTR type). Paraphyses containing a brown vacuolar pigment. Ascospores biguttulate, ornamented with isolated, rounded warts, more rarely with irregular and coalescent warts. Ectal excipulum of textura globulosa with medium-sized to large cells. Species saprobic.

Marcelleina Brumm., Korf & Rifai
Persoonia Suppl., 1: 233  (1967)
Type species: Marcelleina persoonii (P. Crouan & H. Crouan) Brumm. 1967
Family: Pezizaceae
Number of species: 9 fide DoF [4 integrated in 2000 Pezizales]

Mattirolomyces E. Fisch.
Nat. Pflanzenfam. (Ed. 2), 5bVIII: 39  (1938)
Type species: Mattirolomyces terfezioides (Mattir.) E. Fisch. 1938
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Maublancomyces Herter
Revista Südam. de Botanica, 8: 161 (1950)
Type species: Maublancomyces gigas (Krombh.) Herter 1950
Synonym of Gyromitra

Melascypha Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Peziza melaena Fr. 1822
Synonym of Pseudoplectania

Melastiza Boud.
Bull. Soc. mycol. Fr., 1: 106  (1885)
Type species: Melastiza miniata (Fuckel) Boud. 1907
Family: Pyronemataceae
Number of species: 14 fide DoF [5 integrated in 2000 Pezizales]

Microeurotium Ghatak
Ann. Bot. Lond., 50: 860  (1936)
Type species: Microeurotium albidum Ghatak 1936
Family: Incertae sedis
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Microstoma Bernstein
Nova Acta Acad. Caes. Leop.-Carol. Nat. Cur. Dresden, 23: 649  (1852)
Type species: Microstoma hiemale Bernstein 1852
Family: Sarcoscyphaceae
Number of species: 3 fide DoF [3 integrated in 2000 Pezizales]

Midotis Fr.
Elench. fung., 2: 29 (1818)
Type species: Midotis lingua Fr. 1828
Family: Helvellaceae
Number of species: 2 [1 integrated in 2000 Pezizales]

Miladina Svrček
Česká Mykol., 26 (4): 213  (1972)
Type species: Miladina lecithina (Cooke) Svrček 1972
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Asexual morph of Actinosporella type.

Mitrophora Lév.
Ann. Sci. Nat., Bot., sér. 3, 5: 249 (1846)
Type species: Mitrophora semilibera (DC.) Lév. 1846
Synonym of Morchella

Monascella Guarro & Arx
Mycologia, 78 (5): 869 (1986)
Type species: Monascella botryosa Guarro & Arx 1986
Family: Otideaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Moravecia Benkert, Caillet & Moyne
Z. Mykol., 53 (1): 140  (1987)
Type species: Moravecia calospora (Quél.) Benkert, Caillet & Moyne 1987
Family: Pyronemataceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]

Morchella Dill. ex Pers.
Neues Mag. Bot., 1: 116  (1794)
Type species: Morchella esculenta (L.) Pers. 1801
Family: Morchellaceae
Number of species: 36 fide DoF [34 integrated in 2000 Pezizales]

Muciturbo P.H.B. Talbot
Mycol. Res., 92 (1): 95 (1989)
Type species: Muciturbo reticulatus P.H.B. Talbot 1989
Synonym of Ruhlandiella

Mycoarctium K.P. Jain & Cain
Can. J. Bot., 51 (2): 305 (1973)
Type species: Mycoarctium ciliatum K.P. Jain & Cain 1973
Family: Incertae sedis
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Mycoclelandia Trappe & G.W. Beaton
Trans. Br. mycol. Soc., 83 (3): 536  (1984)
Type species: Mycoclelandia arenacea (Trappe) Trappe & G.W. Beaton 1984
Family: Pezizaceae
Number of species: 2 fide DoF [2 integrated in 2000 Pezizales]

Mycogalopsis Gjurašin
Acta bot. Inst. bot. Zagreb, 1: 9  (1925)
Type species: Mycogalopsis retinospora Gjurašin 1925
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Myrmecocystis Harkn.
Proc. Calif. Acad. Sci., Ser. 3, Bot., 1: 269 (1899)
Type species: Myrmecocystis cerebriformis Harkn. 1899
Family: Pyronemataceae
Number of species: 5 [3 integrated in 2000 Pezizales]

Nanoscypha Denison
Mycologia, 64 (3): 617  (1972)
Type species: Nanoscypha tetraspora (Seaver) Denison 1972
Family: Sarcoscyphaceae
Number of species: 10 fide DoF [3 integrated in 2000 Pezizales]
Current definition: Apothecia small, less than 1 cm broad, cup-shaped to discoid or subturbinate, substipitate to stipitate or sessile, centrally attached, less frequently eccentric; texture firm-fleshy to leathery when fresh, brittle and corky when dry; stipe, where present, similar in color and texture to the cup, slender; hymenium concave to discoid, usually some shade of orange but varying from yellow to red, often translucent; exterior whitish or yellowish, glabrous to glabrescent, becoming strongly wrinkled when dry; ectal excipulum sharply differentiated from the medullary excipulum, textura angularis to textura epidermoidea consisting of several layers of large, 5-30 µ, pseudoparenchymatous cells with their long axes perpendicular to the exterior, sometimes with the outermost layer of cells smaller, 1-5 µ, and with thickened walls; medullary excipulum well developed, 40-200 µ or more thick, textura intricata; asci suboperculate with a lateral operculum, iodine negative, less than 300 µ long, maturing seriatim, cylindrical to clavate with a long tapered, and often crooked, base, 4-spored or with a mixture of 8-, 6-, and 4-spored asci; ascospores mostly unequal-sided ellipsoidal (ellipsoidal and symmetrical in N. macrospora), smooth, containing two conspicuous oil drops; paraphyses filiform to subclavate, not anastomosing to form a network about the asci. On leaves, twigs, and small fragments of wood in the tropics.
Asexual morph of Mollardiomyces type.

Napomyces Setch ex Clem. & Shear, , nom. illeg.
Gen. Fungi: 333 (1931)
Type species: Napomyces gardneri (Setch.) Clem. & Shear 1931
Synonym of Daleomyces

Neottiella (Cooke) Sacc.
Syll. fung., 8: 190  (1889)
Type species: Neottiella albocincta (Berk. & M.A. Curtis) Sacc. 1889
Family: Pyronemataceae
Number of species: 13 fide DoF [5 integrated in 2000 Pezizales]

Neournula Paden & Tylutki
Mycologia, 60 (6): 1160  (1969)
Type species: Neournula nordmanensis Paden & Tylutki 1969
Diagnosis: Apothecia stipitata, urceolata vel cupularia; margo crenata vel stellata; hymenium pallidum vel purpureum; externa pallida subbrunnea vel brunnea-purpurea; excipulum medullare e textura intricata; excipulum exterius e textura angulari; pili breves; asci suboperculati, cylindracei, in basim obtusi, in iodo non caerulei, non simul maturescentes; opercula eccentrica: ascosporae ellipsoideae vel oblongateae, verrucoseae, hyalineae; paraphyses tenues, ramosae, anastomosantes
Family: Chorioactidaceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Apothecia stipitate, thin-fleshed, leathery, urceolate to goblet-shaped; margin crenate to stellate; hymenium light colored to dull purplish; exterior pale brown to purplish brown; asci suboperculate, cylindrical, with blunt, occasionally lobed bases, connected to the hypothecium by a narrow hypha, not maturing simultaneously, not bluing in iodine; opercula eccentric; ascospores elliptical or oblong, hyaline, at first smooth, becoming verrucose, the ornamentation staining in heated cotton blue; paraphyses narrow, branched, septate, anastomosing ; medullary excipulum of textura intricata, without an evident gelatinous layer; ectal excipulum of textura angularis, the outermost cells giving rise to short, scattered hairs consisting of individual hyphae.

Nothojafnea Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 93  (1968)
Type species: Nothojafnea cryptotricha Rifai 1968
Family: Pyronemataceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]

Ochotrichobolus Kimbr. & Korf
Mycotaxon, 17: 326 (1983)
Type species: Ochotrichobolus polysporus Kimbr. & Korf 1983
Diagnosis: Differing from Lasiobolus by septate hairs, from Trichobolus by operculate asci, and from both genera by having hairs present only at the margin and by forming buttresses at the bases of the hairs.
Family: Ascodesmidaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Ochromitra Velen.
Monogr. Discom. Bohem.: 391 (1934)
Type species: Ochromitra sphaerospora (Peck) Velen. 1934
Synonym of Gyromitra

Octospora Hedw.
Descr. micr.-anal. musc. frond., 2: 4  (1789)
Type species: Octospora leucoloma Hedw. 1789
Family: Pyronemataceae
Number of species: 84 fide DoF [33 integrated in 2000 Pezizales]

Octosporella Döbbeler
Nova Hedwigia, 31 (4): 827  (1980)
Type species: Octosporella jungermanniarum (P. Crouan & H. Crouan) Döbbeler 1980
Family: Pyronemataceae
Number of species: 7 fide DoF [7 integrated in 2000 Pezizales]

Octosporopsis U. Lindem. & M. Vega
Z. Mykol., 80 (2): 566 (2014)
Type species: Octosporopsis nicolai (Maire) U. Lindem., M. Vega & T. Richt. 2014
Family: Pyronemataceae
Number of species: 2 [2 integrated in 2000 Pezizales]
Current definition: Apothecia 2-7 mm in diameter, sessile, at first more or less obconical, later becoming disc-shaped. Hymenium pale yellow, salmon pink to orange. Margin at first slightly bulged, later more or less membranous, with hyaline hairs; the flanks of the apothecium also covered with hairs. Asci 8-spored, operculate, not amyloid, base pleurorhynchous. Ascospores hyaline, broadly ellipsoid to broadly subfusoid, at maturity thin-walled, with minute refractive light brownish droplets, mostly binucleate, later becoming thick-walled and droplets accumulating in the centre. Paraphyses filiform, apically more or less swollen, terminal cell with faintly to distinctly yellowish homogeneous content. Excipulum two-layered: medullary excipulum of textura intricata/subepidermoidea, composed of thin-walled hyaline cells; ectal excipulum as textura globulosa/angularis, composed of hyaline, thick-walled, isodiametric cells. Hairs hyaline, thick-walled, walls one-layered, multi-septate.

Oedocephalum Preuss
Linnaea, 8: 131 (1851)
Type species: Oedocephalum elegans Preuss 1851
Family: Incertae sedis
Number of species: 9 fide DoF [0 integrated in 2000 Pezizales]

Orbicula Cooke
Handb. Brit. Fungi, 2: 926  (1871)
Type species: Orbicula cyclospora Cooke 1871
Family: Pseudombrophilaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Orcadia G.K. Sutherl.
Trans. Br. mycol. Soc., 5 (1): 151  (1915)
Type species: Orcadia ascophylli G.K. Sutherl. 1915
Family: Incertae sedis
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Ostracoderma Fr.
Syst. orb. veg., 1: 150 (1825)
Type species: Ostracoderma institale Fr. 1825
Family: Pezizaceae
Number of species: 3 fide DoF [0 integrated in 2000 Pezizales]

Otidea (Pers.) Bonord.
Handb. Allgem. mykol.: 205  (1851)
Type species: Otidea onotica (Pers.) Fuckel 1870
Family: Otideaceae
Number of species: 23 fide DoF [29 integrated in 2000 Pezizales]

Oviascoma Y.J. Yao & Spooner
Mycol. Res., 100 (1): 102  (1996)
Type species: Oviascoma paludosum (Dennis) Y.J. Yao & Spooner 1996
Diagnosis: Apothecia solitaria vel gregaria, albida, in sicco brunneola, primum cylindrica demum ovoidea vel obovoidea vel globosa. Discus valde convexus, laevis. Receptaculum emarginatum, glabrum, ad basin angustatum. Excipulum e textura globulosa compositum; muri cellularum tenues; cellulae marginem versus elongatiusculae vel clavatae. Asci operculati, cylindrici, I-. Ascosporae unicellulares, vel subglobosae, spinis ornatae. Paraphyses filiformes, septatae.
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Apothecia solitary or gregarious, whitish when fresh, brownish-orange to brown after draying; cylindric at fist, becoming ovoid or obovoid to almost globose. Disc strongly convex, smooth. Receptacle emarginate, deeply cupulate, outer surface glabrous, attached to the substratum at a narrow base. Excipulum a textura globulosa, cells thin-walied, broadly ellipsoid to subglobose, marginal cells slightly elongate or clavate. Asci operculate, cylindric, I-. Ascospores unicellular, colourless, globose to subglobose, ornamented with spines. Paraphyses filiform, flexuous.

Pachyella Boud.
Hist. Class. Discom. Eur.: 50  (1907)
Type species: Pachyella barlaeana (Bres.) Boud. 1907
Family: Pezizaceae
Number of species: 19 fide DoF [4 integrated in 2000 Pezizales]

Pachyphlodes Zobel
Icon. fung., 6: 55 (1854)
Type species: Pachyphlodes ligerica (Tul. & C. Tul. ex Berk.) Zobel 1854
Family: Pezizaceae
Number of species: 11 fide DoF [8 integrated in 2000 Pezizales]

Pachyphloeus Tul. & C. Tul., nom. illeg.
G. bot. ital., 1 (7-8): 60  (1845)
Type species: Pachyphloeus melanoxanthus (Tul. & C. Tul. ex Berk.) Tul. & C. Tul. 1845
Synonym of Pachyphlodes

Paradiscina Benedix
Kulturpflanze, 17: 274 (1969)
Type species: Paradiscina melaleuca (Bres.) Benedix 1969
Synonym of Gyromitra

Paradoxa Mattir.
Beitr. Kryptfl. Schweiz, 8 (2): 32  (1935)
Type species: Paradoxa monospora Mattir. 1935
Family: Tuberaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Paragalactinia Van Vooren
Ascomycete.org, 12 (4): 188 (2020)
Type species: Paragalactinia succosa (Berk.) Van Vooren 2020
Family: Pezizaceae
Number of species: 10 [7 integrated in 2000 Pezizales]
Current definition: Ascomata hypogeous or epigeous; apothecial species sessile and cupulate. Flesh producing watery latex, becoming yellowsih, greenish or bluish in air. Asci operculate, 8-spored, mainly without crozier, wall weakly bluing in contact with an iodine solution, a little more intensely at the top (WT type). Paraphyses containing a brownish vacuolar pigment. Medullary excipulum containing laticiferous hyphae. Ectal excipulum of textura globulosa, with large cells. Ascospores uni- or biguttulate, warted. Species forming ectomycorrhizas.

Paramitra Benedix
Kulturpflanze, 10: 368 (1962)
Type species: Paramitra ochreoides Benedix 1962
Synonym of Peziza

Parascutellinia Svrček
Česká Mykol., 29: 129  (1975)
Type species: Parascutellinia violacea (Velen.) Svrček 1975
Family: Pyronemataceae
Number of species: 3 fide DoF [1 integrated in 2000 Pezizales]

Paratricharina Van Vooren, U. Lindem., M. Vega, Ribes, Illescas & Matočec
Ascomycete.org, 7 (3): 110 (2015)
Type species: Paratricharina poiraultii (Boud.) Van Vooren, U. Lindemann, M. Vega, Ribes, Illescas & Matočec 2015
Family: Pyronemataceae
Number of species: 1 [3 integrated in 2000 Pezizales]
Current definition: Apothecia about 10–20 mm in diameter or less, sessile, slightly cupulate with inrolled margin, expanded when old; outer surface covered by dense brown-red hairs, sometimes appressed and/or forming distinct pustules, but mostly straight and projecting outwards. Subhymenium thin, of textura intricata. Medullary excipulum of textura intricata, with thin-walled hyphae. Ectal excipulum of vertically oriented textura prismatica, with thin-walled cells, becoming a textura globulosa/subangularis in the outermost part, then with thick-walled brownish cells. Hairs more or less thick-walled, pale brown, more or less sinuous, septate, obtuse or pointed, with a simple base arising from the globose cells of the outer layer of the ectal excipulum. Asci cylindrical, with forked base, arising from perforated croziers, inamyloid, 8-spored. Paraphyses cylindrical, rounded and slightly enlarged at the apex, with spumous compound KOH soluble content, without reaction in Lugol’s solution. Ascospores ellipsoid, hyaline, appearing eguttulate (in dead state) but with small polar aggregations of minute lipid bodies rapidly coalesced into two small polar guttules (in living state), more or less thick-walled, smooth observed in water and Cotton Blue, but with a rough to finely verrucose perispore if mounted in Lugol’s solution, entirely encapsulated by a delicate persistent sheath. Base of apothecia covered by hyaline anchoring hyphae.

Paratrichophaea Trigaux
Docum. Mycol., 16 (61): 5  (1985)
Type species: Paratrichophaea macrocystis Trigaux 1985
Synonym of Lasiobolidium

Parawilcoxina Van Vooren
Ascomycete.org, 13 (1): 36 (2021)
Type species: Parawilcoxina inexpectata Valencia, Van Vooren & M. Vega 2021
Diagnosis: Differs from Wilcoxina by the presence of vacuolar bodies in paraphyses, its saprobic status, and its genetic profile.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, slightly cupuliform, disciform to discoid, whitish to pale greyish, with an external surface covered by small brown hairs. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura globulosa/angularis. Hairs superficial, straight, septate, with a simple base, sometimes arising from a chain of rounded cells. Ascospores uniseriate or irregularly biseriate, ellipsoid to narrow ellipsoid, hyaline, guttulate, smooth. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored. Paraphyses containing vacuolar bodies. Species saprobic. Asexual morph unknown.

Paurocotylis Berk.
Bot. Antarct. Voy. Erebus Terror, 1839-1843 I, Part II, 2: 188  (1855)
Type species: Paurocotylis pila Berk. 1855
Family: Tarzettaceae
Number of species: 1 fide DoF [5 integrated in 2000 Pezizales]

Perilachnea Van Vooren
Ascomycete.org, 13 (1): 18 (2021)
Type species: Perilachnea hemisphaerioides (Mouton) Van Vooren 2021
Diagnosis: Differs from Trichophaea s. str. by deeply cupulate apothecia (at least in young state), mainly with a Humaria-like appearance, bi- or pluriguttulate ascospores, paraphyses containing small oil bodies, a saprobic status, and genetic profile.
Family: Pyronemataceae
Number of species: 3 [6 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, deeply cupuliform or discoid, whitish, yellowish or orange, with an external surface covered by sparse brown hairs. Margin hairy. Excipulum two-layered: medullary layer of textura intricata and ectal layer of textura globulosa/angularis. Hairs superficial, septate, with a simple base. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored. Paraphyses slender, containing small oil bodies. Ascospores uniseriate, ellipsoid, hyaline, bi- or pluriguttulate, smooth or very finely dotted. Species saprobic of the litter and woody debris of conifers or on enriched or burnt soil. Asexual morph unknown but germinating ascospores observed in P. hemisphaerioides and P. flavobrunnea.

Petchiomyces E. Fisch. & Mattir.
Nat. Pflanzenfam. (Ed. 2), 5bVIII: 40  (1938)
Type species: Petchiomyces thwaitesii (Berk. & Broome) E. Fisch. & Mattir. 1938
Family: Pyronemataceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]

Peziza Dill. ex Fr.
Syst. mycol., 2 (1): 40  (1822)
Type species: Peziza vesiculosa Bull. 1790
Family: Pezizaceae
Number of species: 104 fide DoF [28 integrated in 2000 Pezizales]
Asexual morph of Chromelosporium, Oedocephalum type.

Pfistera Korf & W.Y. Zhuang
Mycotaxon, 40: 275 (1991)
Type species: Pfistera pyrophila Korf & W.Y. Zhuang 1991
Synonym of Peziza

Phaeangium Pat.
J. Bot. Paris, 8: 155  (1894)
Type species: Phaeangium lefebvrei Pat. 1894
Synonym of Picoa

Phaeopezia (Vido) Vido
Michelia, 1 (5): 595 (1879)
Type species: Phaeopezia apiculata (Cooke) Vido 1879
Family: Pezizaceae
Number of species: 1 [4 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous or hypogeous; apothecial species sessile and discoid. Asci operculate, 8-spored, with crozier, wall diffusely bluing in an iodine solution (WT type). Paraphyses diffusely coloured (olivaceous brown), with also a brownish external pigment on the top article. Ascospores containing numerous small polar granules, ornamented with isolated warts and with polar apiculi, hyaline becoming brown with age. Excipulum of textura globulosa, with small to medium-sized cells. Species forming ectomycorrhizas.

Phillipsia Berk.
J. Linn. Soc. Bot., 18: 388  (1881)
Type species: Phillipsia domingensis Berk. 1881
Family: Sarcoscyphaceae
Number of species: 17 fide DoF [5 integrated in 2000 Pezizales]

Phylloscypha Van Vooren
Ascomycete.org, 12 (4): 189 (2020)
Type species: Phylloscypha phyllogena (Cooke) Van Vooren 2020
Family: Pezizaceae
Number of species: 5 [5 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, cupulate, sessile, external surface distinctly furfuraceous or pustulate. Flesh without latex, purplish-coloured. Asci operculate, 8-spored, with crozier, wall diffusely bluing in an iodine solution (W type). Paraphyses hyaline, with an external pigment at the top, pale brown, olivaceous brown or dark brown. Ascospores eguttulate but containing small polar granules, ornamented with warts. Species saprobic.

Phymatotrichopsis Hennebert
Persoonia, 7 (2): 199  (1973)
Type species: Phymatotrichopsis omnivora (Duggar) Hennebert 1973
Family: Rhizinaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Physomitra Boud.
Bull. Soc. mycol. Fr., 1: 99 (1885)
Type species: Physomitra esculenta (Pers.) Boud. 1904
Synonym of Gyromitra

Picoa Vittad.
Monogr. Tuberac.: 54  (1831)
Type species: Picoa juniperi Vittad. 1831
Family: Helvellaceae
Number of species: 4 fide DoF [2 integrated in 2000 Pezizales]

Pindara Velen.
Monogr. Discom. Bohem.: 341 (1934)
Type species: Pindara terrestris Velen. 1934
Family: Helvellaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Pithya Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 317  (1870)
Type species: Pithya cupressina (Batsch) Fuckel 1870
Family: Sarcoscyphaceae
Number of species: 5 fide DoF [2 integrated in 2000 Pezizales]
Asexual morph of Mollardiomyces type.

Planamyces Crous & Decock
Persoonia, 39: 377 (2017)
Type species: Planamyces parisiensis Crous & C. Decock 2017
Family: Otideaceae
Number of species: 1 [0 integrated in 2000 Pezizales]

Plectania Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 323  (1870)
Type species: Plectania melastoma (Sowerby) Fuckel 1870
Family: Sarcosomataceae
Number of species: 15 fide DoF [7 integrated in 2000 Pezizales]
Asexual morph of Conoplea type.

Plicaria Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 325  (1870)
Type species: Plicaria trachycarpa (Curr.) Boud. 1907
Family: Pezizaceae
Number of species: 10 fide DoF [4 integrated in 2000 Pezizales]
Asexual morph of Chromelosporium type.

Plicariella (Sacc.) Rehm
Rabenh. Krypt.-Fl. (ed. 2), 1 (3), lief. 42: 988 (1894)
Type species: Plicariella radula (Berk. & Broome) Rehm 1894
Family: Pezizaceae
Number of species: 3 [2 integrated in 2000 Pezizales]

Pseudaleuria Lusk
Mycotaxon, 30: 419  (1987)
Type species: Pseudaleuria quinaultiana Lusk 1987
Diagnosis: Apothecia sparsa vel fasciculata, mediocria, sessilia vel substipitata, orbicularia vel repanda; hymenio clare rubro-aurantiaco, extus similari vel pallidiore, pilis hyalinis, flexuosis, stratum externum velutinum tormantibus; excipulo duobus stratis composito, strato interiore ex textura intricata, strato exteriore ex textura angularis, ascis operculatis, non amyloideis, octosporis, cylindraceis, basin versus sensim angustatis, ad maturitatem non prodtrudentibus, muro tenui; ascosporis ellipsoideis, laevibus, hyalinis, eguttulatis, mediocribus; paraphysibus gracilibus, sursum plus minsve dilatatis, raro ramificantibus.
Family: Pyronemataceae
Number of species: 1 fide DoF [2 integrated in 2000 Pezizales]
Current definition: Apothecia scattered or in clusters, medium sized, sessile to substipitate, discoid to repand, thick-fleshed, texture firm fleshy to rubbery, becoming corky when dry; hymenium bright reddish orange; exterior concolor or lighter when fresh, with flexuous hyphoid hairs, these hairs forming a felty layer; excipulum two-layered, medullary layer is textura intricata, ectal layer is textura angularis; asci operculate, non-amyloid, eight-spored, cylindrical, tapering to the base, thin-walled, not protruding at maturity; ascospores ellipsoid, smooth, hyaline, eguttulate, uninucleate, medium sized; paraphyses slender, slightly to greatly and abruptly enlarged above, rarely branching.Soil and/or wood saprobe, or mycorrhizal.

Pseudoboubovia U. Lindem., M. Vega, B. Perić & R. Tena
Z. Mykol., 81 (2): 391 (2015)
Type species: Pseudoboubovia benkertii (B. Perić) U. Lindem., M. Vega, B. Perić & Tena 2015
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Apothecia 1-8 mm in diameter, sessile, smooth, at first hemispherical, later becoming disc-shaped, finally with irregular shape. Hymenium lemon yellow, greenish yellow, orange yellow. Margin slightly raised, smooth or finely serrated, sometimes irregularly wavy or incurved, same color as the hymenium. Asci 8-spored, operculate, not amyloid, base simple. Ascospores ellipsoid-ovoid or rhomboid, smooth, hyaline or occasionally pale yellowish, with many minute refractive droplets, later becoming thick-walled and droplets accumulating in the centre, with a cyanophilic perispore, dead ascospores containing occasionally a de-Bary-bubble. Paraphyses filiform, distinctly curved, septate, containing yellow or orange granules which do not stain green with iodine. Excipulum two-layered: medullary excipulum of textura intricata, composed of thin-walled hyaline cells; ectal excipulum as textura globulosa, composed of thin walled hyaline cells, intermixed with swollen hyphae. Anchor hyphae covering the base of apothecia.

Pseudocoprotus U. Lindem., Fellm. & J.A. Castillo
Ascomycete.org, 11 (1): 20 (2019)
Type species: Pseudocoprotus catenipilus (J. Moravec) U. Lindem., Fellm. & J.A. Castillo 2019
Diagnosis: Differs from the species of Coprotus sensu Korf & Kimbrough by its marginal hairs composed of globose and elongate cells in chains, its fully developed marginal tissue, its much bigger size, and the rDNA analysis.
Family: Ascodesmidaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Pseudombrophila Boud.
Bull. Soc. mycol. Fr., 1: 106  (1885)
Type species: Pseudombrophila pedrottii Boud. 1885
Family: Pseudombrophilaceae
Number of species: 28 fide DoF [4 integrated in 2000 Pezizales]

Pseudopithyella Seaver
N. Amer. Cup-fungi (Opercul.): 153  (1928)
Type species: Pseudopithyella minuscula (Boud. & Torrend) Seaver 1928
Family: Sarcoscyphaceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]

Pseudoplectania Fuckel
Jahrb. nassau. Ver. Naturk., 23-24: 324  (1870)
Type species: Pseudoplectania nigrella (Pers.) Fuckel 1870
Family: Sarcosomataceae
Number of species: 4 fide DoF [9 integrated in 2000 Pezizales]

Pseudorhizina Jacz.
Opredelitel’ Gribov., 1. Sovershennye Griby: 414  (1913)
Type species: Pseudorhizina korshinskii Jacz. 1913
Family: Discinaceae
Number of species: 2 fide DoF [1 integrated in 2000 Pezizales]

Pseudosarcosoma M. Carbone, Agnello & P. Alvarado
Ascomycete.org, 5 (1): 6 (2013)
Type species: Pseudosarcosoma latahense (Paden & Tylutki) M. Carbone, Agnello & P. Alvarado 2013
Family: Chorioactidaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Pseudotis (Boud.) Boud.
Hist. Class. Discom. Eur.: 52 (1907)
Type species: Pseudotis abietina (Pers.) Boud. 1907
Synonym of Otidea

Pseudotricharina Van Vooren, Tello & M. Vega
Ascomycete.org, 11 (2): 32 (2019)
Type species: Pseudotricharina intermedia Van Vooren, Tello & M. Vega 2019
Diagnosis: Differs from the known species of Tricharina by its guttulate and roughly warted ascospores, and from Trichophaea species by its tricharinoid general morphology.
Family: Pyronemataceae
Number of species: 3 [3 integrated in 2000 Pezizales]

Psilopezia Berk.
London J. Bot., 6: 325  (1847)
Type species: Psilopezia nummularia Berk. 1847
Family: Rhizinaceae
Number of species: 5 fide DoF [1 integrated in 2000 Pezizales]

Ptychoverpa Boud.
Hist. Class. Discom. Eur.: 34 (1907)
Type species: Ptychoverpa bohemica (Krombh.) Boud. 1907
Synonym of Verpa

Pulparia P. Karst.
Bidr. Känn. Finl. Nat. Folk., 19: 9 (1871)
Type species: Pulparia arctica P. Karst. 1871
Synonym of Pulvinula

Pulvinula Boud.
Bull. Soc. mycol. Fr., 1: 107  (1885)
Type species: Pulvinula convexella (P. Karst.) Pfister 1976
Family: Pulvinulaceae
Number of species: 27 fide DoF [4 integrated in 2000 Pezizales]

Purpureodiscus (Hirsch) Van Vooren
Ascomycete.org, 12 (4): 189 (2020)
Type species: Purpureodiscus subisabellinus (Le Gal) Van Vooren 2020
Family: Pezizaceae
Number of species: 4 [2 integrated in 2000 Pezizales]
Current definition: Ascomata discoid or pulvinate, sessile, fleshy, whitish, pinkish to reddish-colored or purplish brown. Flesh without latex. Asci operculate, 8-spored, with crozier, with wall diffusely bluing in an iodine solution (W type). Paraphyses containing small vacuolar bodies. Ascospores eguttulate, but often with small granules at young stage, smooth or ornamented by isolated warts, in some species germinating to directly produce conidia. Ectal excipulum of textura globulosa/angularis, with large cells. Species saprobic.

Pustularia Fuckel, nom. illeg.
Jahrb. nassau. Ver. Naturk., 23-24: 328 (1870)
Type species: Pustularia cupularis (L.) Fuckel 1870
Synonym of Tarzetta

Pustulina Eckblad
Nytt Mag. Bot., 15(1-2): 84 (1968)
Type species: Pustulina cupularis (L.) Eckblad 1968
Synonym of Tarzetta

Pyronema Carus
Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur., 17 (1): 370  (1835)
Type species: Pyronema marianum Carus 1835
Family: Pyronemataceae
Number of species: 2 fide DoF [2 integrated in 2000 Pezizales]

Pyropyxis Egger
Can. J. Bot., 62 (4): 705  (1984)
Type species: Pyropyxis rubra (Peck) Egger 1984
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Asexual morph of Dichobotrys type.

Ramsbottomia W.D. Buckley
Trans. Br. mycol. Soc., 9 (1): 44  (1924)
Type species: Ramsbottomia lamprosporoidea W.D. Buckley 1924
Family: Pyronemataceae
Number of species: 3 fide DoF [1 integrated in 2000 Pezizales]

Reddellomyces Trappe, Castellano & Malajczuk
Aust. Syst. Bot., 5 (5): 606  (1992)
Type species: Reddellomyces westraliensis (G.W. Beaton & Malajczuk) Trappe, Castellano & Malajczuk 1992
Diagnosis: A Labyrinthomyces e Dingleya peridio laevi, glabro et ascis sporis 1–4 (–5) differt.
Family: Tuberaceae
Number of species: 4 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Differing from Labyrinthomyces and Dingleya in its smooth, glabrous peridium and asci with only 1–4 (–5) spores. Ornamentation of spores of some species collections swells inordinately in KOH. True ornamentation can be viewed in H20, Melzer''s reagent or cotton blue.

Rhizina Fr.
Observ. mycol., 1: 161  (1815)
Type species: Rhizina undulata Fr. 1815
Family: Rhizinaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Rhizoblepharia Rifai
Verh. K. ned. Akad. Wet., tweede sect., 57 (3): 104  (1968)
Type species: Rhizoblepharia jugispora Rifai 1968
Family: Pyronemataceae
Number of species: 2 fide DoF [0 integrated in 2000 Pezizales]

Rhizopodella (Cooke) Boud.
Bull. Soc. mycol. Fr., 1: 103 (1885)
Type species: Peziza melastoma Sowerby 1798
Synonym of Plectania

Rhodopeziza Hohmeyer & J. Moravec
Czech Mycol., 47 (4): 261  (1995)
Type species: Rhodopeziza tuberculata (Gamundí) J. Moravec & Hohmeyer 1995
Diagnosis: Apothecia media usque magna, sessilia, cupuliformia, inaequalia, interdum auriculiformia, externe glabra, ad marginem pruinosa; hymenio miniato. Excipulum externum textura globulosa angularis compositum e cellulis latioribus, excipulum internum (medulla) textura globulosa angularis compositum e cellulis angustioribus cum hyphis intermixtis. Asci cylindracei, operculati, octospori, in toto iodo coerulescentes. Ascosporae late ellipsoideae, eguttulatae, pallide flavescentes, tuberculatae. Paraphyses filiformes, simplices, septatae, rectae sed apice curvatae, granulis aurantiacis impletae.
Family: Pezizaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Rhodoscypha Dissing & Sivertsen
Mycotaxon, 16 (2): 442  (1983)
Type species: Rhodoscypha ovilla (Peck) Dissing & Sivertsen 1983
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Rhodotarzetta Dissing & Sivertsen
Mycotaxon, 16 (2): 453  (1983)
Type species: Rhodotarzetta rosea (Rea) Dissing & Sivertsen 1983
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Rickiella Syd. & P. Syd. ex Rick
Ann. mycol., 2 (3): 244  (1904)
Type species: Rickiella transiens Syd. & P. Syd. ex Rick 1904
Family: Sarcoscyphaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Ruhlandiella Henn.
Hedwigia, 42 (Beibl.): 24  (1903)
Type species: Ruhlandiella berolinensis Henn. 1903
Family: Pezizaceae
Number of species: 1 fide DoF [5 integrated in 2000 Pezizales]
Current definition: Fruitbodies pulvinate to brain-like. Excipulum of globose to angular, thick-walled cells. Asci cylindrical to clavate, weakly amyloid all over, without operculum; ascus wall evanescent. Paraphyses septate, branching, with a conspicuous gelatinous sheath; paraphyses distinctly exceeding asci in length. Ascospores globose, at first bi- to tri-seriate, hyaline, later uniseriate, with a pale brownish, strongly cyanophilic ornamentation.

Saccobolus Boud.
Ann. Sci. Nat. Bot., sér. 5, 10: 228  (1869)
Type species: Saccobolus kervernii (P. Crouan & H. Crouan) Boud. 1907
Family: Ascobolaceae
Number of species: 27 fide DoF [15 integrated in 2000 Pezizales]

Sarcopeziza Loizides, Agnello & P. Alvarado
Ascomycete.org, 10 (4): 179 (2018)
Type species: Sarcopeziza sicula (Inzenga) Agnello, Loizides & P. Alvarado 2018
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Ascomata hypogeous or semi-hypogeous, medium- to large-sized, sarcosphaeroid in aspect and distinctly or indistinctly stipitate, at first angiocarpic, hollow, subglobose to pyriform, with a small apical orifice, gradually tearing concentrically to form erect or arched lacinias; hymenial surface smooth, purplish-violet; asci with fully functioning opercula, , pleurorhynchous, diffusely amyloid over the entire length, more intensely at the apex; paraphyses multiseptate, cylindrical or sometimes moniliform; ascospores monoseriate, ellipsoid, smooth or finely low warted, hyaline or subhyaline and mono- or multiguttulate,; in xeric scrub, dry grasslands and thermomediterranean matorral.

Sarcoscypha (Fr.) Boud.
Bull. Soc. mycol. Fr., 1: 103  (1885)
Type species: Sarcoscypha coccinea (Jacq.) Sacc. 1889
Family: Sarcoscyphaceae
Number of species: 28 fide DoF [14 integrated in 2000 Pezizales]
Asexual morph of Mollardiomyces type.

Sarcosoma Casp.
Rabenh. Krypt.-Fl. Edn, 2, 1.3: 497  (1891)
Type species: Sarcosoma globosum (Schmidel) Casp. 1891
Family: Sarcosomataceae
Number of species: 5 fide DoF [1 integrated in 2000 Pezizales]
Asexual morph of Verticicladium type.

Sarcosphaera Auersw.
Hedwigia, 8: 82  (1869)
Type species: Sarcosphaera macrocalyx (Riess) Auersw. 1869
Family: Pezizaceae
Number of species: 1 fide DoF [2 integrated in 2000 Pezizales]

Scabropezia Dissing & Pfister
Nord. J. Bot., 1 (1): 102  (1981)
Type species: Scabropezia scabrosa (Cooke) Dissing & Pfister 1981
Synonym of Plicariella

Scodellina Gray
Nat. arrang. Brit. plants, 1: 669 (1821)
Type species: Scodellina vesiculosa (Bull.) Gray 1821
Synonym of Peziza

Scutellinia (Cooke) Lambotte
Mém. Soc. roy. Sci. Liège, sér. 2, 14: 299  (1887)
Type species: Peziza scutellata L. 1753
Family: Pyronemataceae
Number of species: 66 fide DoF [26 integrated in 2000 Pezizales]

Selenaspora R. Heim & Le Gal
Rev. Mycol. Paris, 1: 307  (1936)
Type species: Selenaspora batava R. Heim & Le Gal 1936
Family: Sarcosomataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Sepultaria (Cooke) Boud.
Bull. Soc. mycol. Fr., 1: 104  (1885)
Type species: Peziza sepulta Fr. 1851
Family: Pyronemataceae
Number of species: unknown

Sepultariella Van Vooren, U. Lindem. & Healy
Ascomycete.org, 9 (4): 118 (2017)
Type species: Sepultariella patavina (Cooke & Sacc.) Van Vooren, U. Lindemann & Healy 2017
Diagnosis: Differs from Leucoscypha Boud. by pigmented apothecia and smooth ascospores.
Family: Pyronemataceae
Number of species: 1 fide DoF [2 integrated in 2000 Pezizales]

Smardaea Svrček
Česká Mykol., 23: 90  (1969)
Type species: Smardaea amethystina (W. Phillips) Svrček 1969
Family: Pyronemataceae
Number of species: 7 fide DoF [9 integrated in 2000 Pezizales]

Smarodsia Raitv. & Vimba
Folia cryptog. Estonica, 42: 98 (2006)
Type species: Smarodsia stollii Raitv. & Vimba 2006
Diagnosis: Genus pyronematacearum Kotlabaea Svrcek similis sed in sporis distincte guttulatis et pilis setiformibus crassiter tunicatis differt.
Family: Pyronemataceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]
Current definition: Apothecia terrestrial, minute, broadly sessile, discoid to saucer-shaped, disc pale yellowish to pale ochraceous or orange-yellowish, receptacle concolorous with the disc, externally smooth. Ectal excipulum of hyaline, large-celled textura globulosa. The cells of outer layers have thick, refracting walls. Hairs scanty, hyaline, aseptate, firm-walled to thick-walled, cylindrical to conical, obtusely rounded, rarely almost pointed, straight to slightly flexuous, arising superficially from a basal cell. Asci pleurorhynchous, cylindrical, apically not bluing in MLZ, at first 8-spored but 4-spored at maturity in type species. Ascospores ellipsoid-fusoid, containing 2 big, more rarely 1 big or 1 big and several small lipid globules, sometimes with numerous small globules or granulose cytoplasma, spore wall without changes when heated in lactic acid. Paraphyses apically clavate, fi lled with numerous small yellowish drops in CB.

Sowerbyella Nannf.
Svensk bot. Tidskr., 32: 118  (1938)
Type species: Sowerbyella radiculata (Sowerby) Nannf. 1938
Family: Pyronemataceae
Number of species: 16 fide DoF [9 integrated in 2000 Pezizales]

Spaniodiscus Van Vooren, U. Lindemann & Roffler
Ascomycete.org, 14 (2): 43 (2022)
Type species: Spaniodiscus fuckelii (Donadini & Svrček) Van Vooren, Roffler & U. Lindem. 2022
Diagnosis: Differs from Scutellinia and other genera having an orange hymenium by the combination of the following characters: superficial and simple-based hairs, always smooth thick-walled ascospores, filled with oil droplets, and its genetic profile.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, orange-coloured, with an external surface and margin covered by short brown hairs. Excipulum two-layered: medullary layer of textura intricata, ectal layer of textura globulosa/angularis. Hairs simple, superficial, septate, with a simple base. Asci operculate, arising from croziers, inamyloid, 8-spored. Paraphyses slender, enlarged at the top, containing carotenoid pigments. Ascospores uniseriate, ellipsoid, hyaline, pluriguttulate in living state, smooth. Trophic status probably saprobic. Asexual morph unknown.

Sphaeridiobolus Boud.
Bull. Soc. mycol. Fr., 1: 108 (1885)
Type species: Ascobolus hyperboreus P. Karst. 1871
Synonym of Ascobolus

Sphaerosoma Klotzsch
Fl. Regn. Boruss., 7: 467  (1839)
Type species: Sphaerosoma fuscescens Klotzsch 1839
Family: Pyronemataceae
Number of species: 3 fide DoF [0 integrated in 2000 Pezizales]

Sphaerosporella (Svrček) Svrček & Kubička
Česká Mykol., 15 (2): 66 (1961)
Type species: Sphaerosporella brunnea (Alb. & Schwein.) Svrček & Kubička 1961
Synonym of Trichophaea

Sphaerozone Zobel
Icon. fung. (Prague), 6: 52  (1854)
Type species: Sphaerozone tulasnei Zobel 1854
Family: Pezizaceae
Number of species: 4 fide DoF [1 integrated in 2000 Pezizales]

Spooneromyces T. Schumach. & J. Moravec
Nord. J. Bot., 9 (4): 426  (1989)
Type species: Spooneromyces laeticolor (P. Karst.) T. Schumach. & J. Moravec 1989
Family: Pyronemataceae
Number of species: 3 fide DoF [3 integrated in 2000 Pezizales]

Stephensia Tul. & C. Tul.
C. r. hebd. Séanc. Acad. Sci. Paris, 21: 1433  (1845)
Type species: Stephensia bombycina (Vittad.) Tul. 1851
Synonym of Hydnocystis

Strobiloscypha N.S. Weber & Denison
Mycotaxon, 54: 130  (1995)
Type species: Strobiloscypha keliae N.S. Weber & Denison 1995
Diagnosis: Apothecia sessilia, juventute aliquantum turbinata, maturitate vade crateriformia vel plana, 1.5-5.0(-7.0) mm in diam, succulenta. Hymenium olivaceo-brunneolum vel olivaceo-cinereum. Pagina abhymenialis brunneola vel brunnea, glabra, margo minute pustullata. Sporae ellipsoideae, (18.7-)21-23.7 x (11.8-)12.5-13(-15.1) µm, hyalinae, maturitatae agullulatae, verruculosae, in ascis uniseriate. Asci J-, operculati, 8-sporae, 316-368 x 14.3-18.4 µm. Paraphyses rectae, anguste clavatae, septatae. Subhymenium e excipulum medullosum texturis intricatis, hyphae 5.3-10.5(-15.6) µm in diam. Excipulum ectale texturis globulosis/angularis. In strobilis Cupressacearum e Taxodiacearum.
Family: Chorioactidaceae
Number of species: 1 fide DoF [2 integrated in 2000 Pezizales]

Tarzetta (Cooke) Lambotte
Mém. Soc. roy. Sci. Liège, sér. 2, 14: 325  (1888)
Type species: Peziza tarzetta Cooke 1877
Family: Tarzettaceae
Number of species: 9 fide DoF [12 integrated in 2000 Pezizales]

Temperantia K. Hansen, Healy & Kovács
Mycologia, 103 (4): 837 (2011)
Type species: Temperantia tiffanyae (Healy) K. Hansen, Healy & Kovács 2011
Family: Pezizaceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Terfezia (Tul. & C. Tul.) Tul. & C. Tul.
Fungi hypog.: 172  (1851)
Type species: Terfezia arenaria (Moris) Trappe 1971
Family: Pezizaceae
Number of species: 12 fide DoF [15 integrated in 2000 Pezizales]

Terracavicola A.C. Grupe, Kraisit., Guevara & M.E. Sm.
Ascomycete.org, 11 (2): 41 (2019)
Type species: Terracavicola echinospora A.C. Grupe, Kraisit., Guevara & M.E. Sm. 2019
Diagnosis: A hypogeous, creamy-tan to light brown truffle-like ascoma with a roughened peridium that is covered with copious brown hairs (setae), the presence of a weak epithecium which covers the hymenium and prevents active discharge of the ascospores, and unique ITS and 28S DNA sequences.
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Thecotheus Boud.
Ann. Sci. Nat. Bot., sér. 5, 10: 235  (1869)
Type species: Thecotheus pelletieri (P. Crouan & H. Crouan) Boud. 1869
Family: Ascobolaceae
Number of species: 17 fide DoF [12 integrated in 2000 Pezizales]

Thindia Korf & Waraitch
Mycologia, 63 (1): 102  (1971)
Type species: Thindia cupressi Korf & Waraitch 1971
Family: Sarcoscyphaceae
Number of species: 1 fide DoF [1 integrated in 2000 Pezizales]

Tirmania Chatin
Truffe (ed. 2): 80  (1892)
Type species: Tirmania nivea (Desf.) Trappe 1971
Family: Pezizaceae
Number of species: 3 fide DoF [2 integrated in 2000 Pezizales]

Trichaleurina Rehm
Ann. mycol., 1 (6): 515 (1903)
Type species: Trichaleurina polytricha Rehm 1914
Family: Chorioactidaceae
Number of species: 4 [2 integrated in 2000 Pezizales]

Tricharia Boud., nom. illeg.
Bull. Soc. mycol. Fr., 1: 104 (1885)
Type species: Tricharia gilva (Cooke) Boud. 1904
Synonym of Tricharina

Tricharina Eckblad
Nytt Mag. Bot., 15 (1-2): 60  (1968)
Type species: Tricharina gilva (Boud. ex Cooke) Eckblad 1968
Family: Pyronemataceae
Number of species: 13 fide DoF [5 integrated in 2000 Pezizales]
Asexual morph of Ascorhizoctonia type.

Tricharinopsis U. Lindem., Van Vooren & Healy
Ascomycete.org, 11 (5): 155 (2019)
Type species: Tricharinopsis herinkii (Svrček) U. Lindem., Van Vooren & Healy 2019
Diagnosis: Differs from Tricharina Eckblad by the absence of clustered hairs at the margin and by an outer surface densely covered with long hyaline to brown flexuous hairs (reminiscent of a Geopora species).
Family: Pyronemataceae
Number of species: 1 [1 integrated in 2000 Pezizales]

Trichobolus (Sacc.) Kimbr. & Cain
Amer. J. Bot., 54 (1): 20 (1967)
Type species: Trichobolus zukalii (Heimerl) Kimbr. 1967
Family: Ascodesmidaceae
Number of species: 4 fide DoF [2 integrated in 2000 Pezizales]

Trichophaea Boud.
Bull. Soc. mycol. Fr., 1: 105  (1885)
Type species: Trichophaea woolhopeia (Cooke & W. Phillips) Arnould 1893
Diagnosis: Poils du réceptacle longs et aigus ; spores le plus souvent garnies de gouttelettes intérieures. Hyménium blanchâtre ou glauque.
Family: Pyronemataceae
Number of species: 25 fide DoF [9 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, subglobose, discoid or cupuliform, hymenium whitish, pale grey to grey-brown, brown or blackish, with an external surface covered by superficial brown hairs. Margin hairy. Excipulum two-layered: medullary layer thick, of textura intricata and ectal layer of textura subglobulosa/angularis. Hairs superficial, septate, with a simple base. Asci operculate, arising from croziers, inamyloid, 8-spored. Paraphyses slender, equal to enlarged (sometimes capitate) at the top, hyaline. Ascospores uniseriate, ellipsoid or spherical, hyaline, uniguttulate, more rarely biguttulate, thick-walled, smooth. Species forming ectomycorrhizae.
Asexual morph of Dichobotrys type.

Trichophaeopsis Korf & Erb
Phytologia, 24 (1): 18  (1972)
Type species: Trichophaeopsis bicuspis (Boud.) Korf & Erb 1972
Family: Pyronemataceae
Number of species: 3 fide DoF [4 integrated in 2000 Pezizales]
Current definition: Ascomata epigeous, apothecial, sessile, pale greyish, with an external surface and margin covered by upward-pointing brown hairs. Excipulum three-layered: medullary layer of textura intricata, ectal layer of textura globulosa/angularis and the outermost part of ectal excipulum of textura prismatica. Hairs simple or bifurcate, deeply inserted in the excipulum, septate, mainly with a simple base. Asci operculate, narrowing toward base, arising from croziers, inamyloid, 8-spored, more rarely 4-spored. Paraphyses slender, not enlarged at the top. Ascospores uniseriate, ellipsoid, hyaline, eguttulate, but sometimes containing small granules that can merge into two oil drops in overmature or dead ascospores, smooth or verrucose. Species saprobic, on soil, woody debris or organic matter. Asexual morph unknown.

Tuber P. Micheli ex F.H. Wigg.
Prim. fl. holsat.: 1-112  (1780)
Type species: Tuber aestivum Vittad. 1831
Family: Tuberaceae
Number of species: 86 fide DoF [38 integrated in 2000 Pezizales]

Tubipeda Falck
Mykol. Untersuch. Ber., 1 (3): 377 (1923)
Type species: Tubipeda elastica (Bull.) Falck 1923
Synonym of Helvella

Ulurua Trappe, Claridge & Kovács
Mycol. Progr., 9 (1): 140 (2010)
Type species: Ulurua nonparaphysata Trappe & Claridge 2010
Diagnosis: Ascomata usque ad 4 cm lata, sulcata, globosa vel turbinata, tomentosa, brunnea. Gleba venis labyrinthinis, vacuis vel hymeniis opponentibus repletis. Sporae hyalinae, late ellipsoideae vel subglobosae, 12–15 x 10–12 μm, verrucis minutis ornatae. Asci in hymenio innati, amyloidei, sporis octo. Paraphyses destitutae. Peridium glebaque cellulis multo inflatis.
Family: Pezizaceae
Number of species: 1 [0 integrated in 2000 Pezizales]

Underwoodia Peck
Ann. Rep. N.Y. St. Mus., 43: 32  (1890)
Type species: Underwoodia columnaris Peck 1890
Family: Helvellaceae
Number of species: 3 fide DoF [1 integrated in 2000 Pezizales]

Urceolaria Bonord., nom. illeg.
Handb. Allgem. mykol.: 203, 311 (1851)
Type species: Urceolaria agelaea (Ach.) Ach. 1803
Family: Incertae sedis
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Urnula Fr.
Summa veg. Scand., section post.: 364  (1849)
Type species: Urnula craterium (Schwein.) Fr. 1851
Family: Sarcosomataceae
Number of species: 6 fide DoF [8 integrated in 2000 Pezizales]

Verpa Sw.
K. svenska Vetensk-Akad. Handl., 36: 129  (1815)
Type species: Verpa conica (O.F. Müll.) Sw. 1815
Family: Morchellaceae
Number of species: 5 fide DoF [2 integrated in 2000 Pezizales]

Verticicladium Preuss
Linnaea, 24: 127 (1851)
Type species: Verticicladium trifidum Preuss 1851
Family: Incertae sedis
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Warcupia Paden & J.V. Cameron
Can. J. Bot., 50 (5): 999  (1972)
Type species: Warcupia terrestris Paden & J.V. Cameron 1972
Family: Otideaceae
Number of species: 1 fide DoF [2 integrated in 2000 Pezizales]

Wenyingia Zheng Wang & Pfister
Mycotaxon, 79: 397 (2001)
Type species: Wenyingia sichuanensis Zheng Wang & Pfister 2001
Diagnosis: Apothecia cupulata, stipitata, brunneloa, usque lO mm diam, 22 mm altitudo. Excipulum ectal e cellulis brunneloa, angularibus 5-20 µm diam (textura angulari), excipulum medullare ex hyphis brunneae. Compositum. Asci octospori, crasse tunicati, cylindracei, 270-295 × 13.5-16 µm. Ascospori ellipsoidei, biguttulati, 19-21.5 × 11-12 µm.
Family: Tarzettaceae
Number of species: 1 fide DoF [0 integrated in 2000 Pezizales]

Wilcoxina Chin S. Yang & Korf
Mycotaxon, 24: 511  (1985)
Type species: Wilcoxina mikolae (Chin S. Yang & H.E. Wilcox) Chin S. Yang & Korf 1985
Diagnosis: Ab Tricharina excipulo ectali non distincte bistrato, ascosporis dimidium superius asci maturi tantum occupantibus, et procreatione anamorphosis lente crescentis generis Complexipedis et non celeriter crescentis generis Ascorhizoctoniae distinguenda.
Family: Pyronemataceae
Number of species: 3 fide DoF [2 integrated in 2000 Pezizales]
Current definition: Apothecia small, cupulate, expanding to discoid, sessile, attached only at the center, scattered or gregarious; hymenium whitish to yellowish orange, senescing to brownish gray; hairs both marginal and running down the flanks, fasciculate, with large clusters near margin and clusters gradually becoming smaller towards the base of the apothecium, pale brown to brown, with narrow bases arising from a chain of thick-walled excipular cells, the broadest part of the hair well above the point of origin of the hair, containing brownish, globose inclusions often appearing attached to the walls, walls 2.0-4.0 µm thick, with thin or thick septa; excipulum clearly two-layered near the margin, ental layer becoming broader toward the apothecial base, of textura intricata, in some taxa having refractive, thick-walled hyphae threading horizontally through the layer, ectal layer of textura globulosa to textura angularis, composed of some almost globose to radially elongated cells with pale brown to brown, thick walls becoming mixed near the apothecial base with some small-diameter hyphae from the ental layer; asci cylindrical, narrowing toward base, apically operculate, 8-spored or rarely (2-) 4 (-6)-spored, J-; ascospores ellipsoid, uniseriate, lining up in the upper half of the ascus, smooth, hyaline or refractive-yellow when mature and mounted in media containing lactic acid, eguttulate, but often with a few small granules or with small, polar clusters of fine granules; paraphyses slender, septate, branched near base, with or without carotenoid particles, slightly enlarged toward apex.
Asexual morph of Complexipes type.

Wolfina Seaver ex Eckblad
Nytt Mag. Bot., 15 (1-2): 126  (1968)
Type species: Wolfina aurantiopsis (Ellis) Seaver 1937
Family: Chorioactidaceae
Number of species: 3 fide DoF [1 integrated in 2000 Pezizales]

Wynnea Berk. & M.A. Curtis
J. Linn. Soc. Bot., 9: 424  (1867)
Type species: Wynnea gigantea Berk. & M.A. Curtis 1867
Family: Wynneaceae
Number of species: 4 fide DoF [3 integrated in 2000 Pezizales]

Wynnella Boud.
Bull. Soc. mycol. Fr., 1: 102 (1885)
Type species: Wynnella auricula (Schaeff.) Boud. 1885
Synonym of Midotis

 

Last Update on 04/25/2022